139736eo.pdf (20MB) - Japan Oceanographic Data Center
139736eo.pdf (20MB) - Japan Oceanographic Data Center 139736eo.pdf (20MB) - Japan Oceanographic Data Center
in 6 months when spawning occurs. They occur where the depth exceeds 150 m, which in the presentcontext would be the shelf edge. Vertical migrations occur between 150 to 200 m depth in the dayrising to within 10 m of the surface at night.Stock densities in the Gulf of Aden were estimated at above 100 tons km-2 (GJOSAETER,1984). Based on growth and recruitment studies, SANDERS and BOUHLEL (1982), estimated theannual yield from the Gulf of Aden at about 15 x lo6 tons.GENERAL CHARACTERISTICS OF EXPLOITABLE RESOURCESDemersal fish populations vary in composition and standing stocks with season, depth andlocality on the coast. In the west around Aden the dominant taxa in the non-upwelling season areLeiognathidae, Carangidae, Pomadasyidae, Lethrinidae and Nemipteridae. During upwellingCentrolophidae, Sparidae, Synodontidae, Sepia sp. and Loligo spp. are prominent. In the east aroundRas Fartak the dominant non-upwelling taxa are Sepia, Ariidae, Sparidae, Dasyatidae andSphyraenidae, while during upwelling Callionymidae, Triglidae, Nemipteridae, Clupeidae andS y nodontidae predominate.Standing stocks at 30 m depth have been measured to be 3.6 and 6.0 tons km-2 in the westand east respectively during non-upwelling, falling to 1.9 and 2.5 tons km-2 during upwelling.Maximum standing stocks occur at 30 m depth in October, at 110 m in May and 20 m or less inAugust.During upwelling demersal populations migrate both inshore and vertically in the water, andappear to separate into pockets along the coast. Deeper water species move on to the shelf and intoshallow water at this time. These migrations probably occur in response to oxygen minima.Pelagic fish stocks are concentrated along the coast in the non-upwelling season at Aden,Mukalla and around Ras Fartak. During upwelling part of the stocks in the east migrate offshore,while those at Aden remain inshore. Pelagic trawls have showed Clupeidae (i.e. Surdinella longiceps ,the oil sardine) to be dominant in the non-upwelling period.Large pelagic fish (e.g. tunas, Spanish mackerels) have not yet been assessed, butconcentrations appear mainly in the western two thirds of the Gulf, with highest stocks in the Adenregion. Here substantial stocks of Indian mackerel, Rastrelliger kanagurta, are found.Total demersal stocks may exceed 116 x lo3 tons, while pelagic stocks may be around 300 x103 tons.SUMMARYThis region of the Indian Ocean presents a rather unique, large-size ecosystem of which themost remarkable characteristics are as follows:Oceanographic conditions that reverse in response to the changes between NE and SWmonsoons also respond to variable and poorly understood influences of the SomaliCurrent and inputs from the Red Sea and the North Arabian Sea intermediate - deep watermasses.Upwelling occurs regularly during the SW monsoon season in many areas of this region.However, the most affected areas are in the east where the upwelling is much greater thanin “classic” upwelling regions of the world oceans.Because of the above conditions, the supply of nutrients to euphotic layers is suffici:ntthroughout the annual cycle to support a balanced primary production. During the summerand the subsequent postmonsoon season, however, the production dramatically increases,reaching densities up to 6 million cells dm-3 and 5 pg dm-3 of chlorophyll a. Secondaryproduction of zooplankton is also very high, with average dry weight biomass of about100 mg m3. This suggests that these waters are among the richest not only within theIndian Ocean but also in the world oceans.269
It is logical to expect that such high productivity supports very large stocks of exploitableresources. This expectation is true for the largely underexploited clupeid, carangid andscombrid fishes and an immense potential (est. 15 million tons) of mesopelagicmyctophids. However, stocks of demersal fish (which are at present the target ofcommercial fisheries) appear to be limited and also heavily exploited.Unfortunately the overall high pelagic productivity in an “immature” upwelling ecosystemindirectly contributes to the disequilibrium of its oxygen balance, as a large part ofproduced organic matter is “wasted” by bacterial decomposition. In the case of the Gulf ofAden this is combined with the general deficit of the oxygen in the North Arabian Sea.Consequently, in the largest part of the Gulf the semi-anoxic conditions (0.5 ml 1-1 or lessof dissolved oxygen) appear throughout the upper water column and below 150 - 200 mdepth. During the summer this anoxia is generally pushed into the surface layers, and canbe extreme in upwelling centres where semi-anoxic conditions prevail below a depth of 20m or even less.Anoxia, which can be considered as an ecological catastrophy, affects the ecosystem inmany poorly understood ways. It certainly contributes to an extraordinarily low standingstock of the benthic communities on the shelf bottoms, and consequently to the missing orvery weak relationship of the benthos with the demersal fish populations. This in turnalso means the trophic dependence of demersal fish on pelagic food potentials that leads togreater degree of overall instability. The appearance of anoxia in shelf waters also causeslarge migrations and structural recombinations in demersal fish communities, and mostlikely the migrations of epipelagic fish also. Although anoxia-induced fishmass-mortalities have not been observed in the Gulf of Aden, “hidden” mortalities areassumed to occur.REGIONAL COOPERATIVE INVESTIGATIONS OF THE NW ARABIAN SEAThe unique oceanographic and marine biological phenomena in the Gulf of Aden and adjacentparts of the Arabian Sea are still poorly documented, and therefore deserve greater attention.Moreover, the crucial economic and nutrition needs of bordering nations the Gulf of Aden providemore practical reasons for increased research. The development of a rational and economically healthyexploitation of the living resources in this region, however, cannot work properly without an adequatescientific basis. To achieve such a scientific basis requires a rather complex knowledge on theecosystem and living resources (Fig. 19). Using BEHRMANs (1981) words, “finding fish is notenough”; it leads to the tasks as shown in the lower part of Figure 19.Since the Gulf of Aden and adjacent parts of the Arabian Sea inevitably must be considered asa functional unit, at least from the standpoint of bioproductivity and oceanographic processes,investigations must cover the whole region. However, the area is extensive and the seas, particularlyduring the summer monsoon, are heavy. Therefore the sea-going research requires a sizeable wellequippedresearch vessel, operating continuously year around with an interdisciplinary scientific teamaboard, covering the area as shown in Figure 20. Probably no single country bordering this area(Djibouti, Somalia, P.D.R. Yemen, A.R. Yemen and Oman) alone can support the costs of such astudy, and even so, this would lead to an uneconomic duplication of efforts. Therefore it is proposedherewith to set up a regionally operating organization, shared by all bordering countries and supportedby an UN interagency group (UNDP/FAO/UNESCO-IOC/UNEP). A tentative outline of scientificprogramme of interdisciplinary oceanographic and fisheries investigations for this area is proposed inthe Appendix.270
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It is logical to expect that such high productivity supports very large stocks of exploitableresources. This expectation is true for the largely underexploited clupeid, carangid andscombrid fishes and an immense potential (est. 15 million tons) of mesopelagicmyctophids. However, stocks of demersal fish (which are at present the target ofcommercial fisheries) appear to be limited and also heavily exploited.Unfortunately the overall high pelagic productivity in an “immature” upwelling ecosystemindirectly contributes to the disequilibrium of its oxygen balance, as a large part ofproduced organic matter is “wasted” by bacterial decomposition. In the case of the Gulf ofAden this is combined with the general deficit of the oxygen in the North Arabian Sea.Consequently, in the largest part of the Gulf the semi-anoxic conditions (0.5 ml 1-1 or lessof dissolved oxygen) appear throughout the upper water column and below 150 - 200 mdepth. During the summer this anoxia is generally pushed into the surface layers, and canbe extreme in upwelling centres where semi-anoxic conditions prevail below a depth of 20m or even less.Anoxia, which can be considered as an ecological catastrophy, affects the ecosystem inmany poorly understood ways. It certainly contributes to an extraordinarily low standingstock of the benthic communities on the shelf bottoms, and consequently to the missing orvery weak relationship of the benthos with the demersal fish populations. This in turnalso means the trophic dependence of demersal fish on pelagic food potentials that leads togreater degree of overall instability. The appearance of anoxia in shelf waters also causeslarge migrations and structural recombinations in demersal fish communities, and mostlikely the migrations of epipelagic fish also. Although anoxia-induced fishmass-mortalities have not been observed in the Gulf of Aden, “hidden” mortalities areassumed to occur.REGIONAL COOPERATIVE INVESTIGATIONS OF THE NW ARABIAN SEAThe unique oceanographic and marine biological phenomena in the Gulf of Aden and adjacentparts of the Arabian Sea are still poorly documented, and therefore deserve greater attention.Moreover, the crucial economic and nutrition needs of bordering nations the Gulf of Aden providemore practical reasons for increased research. The development of a rational and economically healthyexploitation of the living resources in this region, however, cannot work properly without an adequatescientific basis. To achieve such a scientific basis requires a rather complex knowledge on theecosystem and living resources (Fig. 19). Using BEHRMANs (1981) words, “finding fish is notenough”; it leads to the tasks as shown in the lower part of Figure 19.Since the Gulf of Aden and adjacent parts of the Arabian Sea inevitably must be considered asa functional unit, at least from the standpoint of bioproductivity and oceanographic processes,investigations must cover the whole region. However, the area is extensive and the seas, particularlyduring the summer monsoon, are heavy. Therefore the sea-going research requires a sizeable wellequippedresearch vessel, operating continuously year around with an interdisciplinary scientific teamaboard, covering the area as shown in Figure 20. Probably no single country bordering this area(Djibouti, Somalia, P.D.R. Yemen, A.R. Yemen and Oman) alone can support the costs of such astudy, and even so, this would lead to an uneconomic duplication of efforts. Therefore it is proposedherewith to set up a regionally operating organization, shared by all bordering countries and supportedby an UN interagency group (UNDP/FAO/UNESCO-IOC/UNEP). A tentative outline of scientificprogramme of interdisciplinary oceanographic and fisheries investigations for this area is proposed inthe Appendix.270