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The Malay archipelago : the land of the orang-utan ... - Wallace Online

The Malay archipelago : the land of the orang-utan ... - Wallace Online

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XVIII.] NATURAL HISTORY OF CELEBES. 215<strong>The</strong>se large arid well-known families well represent <strong>the</strong>general character <strong>of</strong> <strong>the</strong> zoology <strong>of</strong> Celebes ; and <strong>the</strong>y showthat this is<strong>land</strong> is really one <strong>of</strong> <strong>the</strong> most isolated joortions <strong>of</strong><strong>the</strong> Archipelago, although situated in its very centre.But <strong>the</strong> insects <strong>of</strong> Celebes present us with o<strong>the</strong>r phenomenamore curious and more difficult to explain than <strong>the</strong>ir specificindividuality. <strong>The</strong> butterflies <strong>of</strong> that is<strong>land</strong> ai'e in many casescharacterized by a peculiarity <strong>of</strong> outline, which distinguishes<strong>the</strong>m at a glance from those <strong>of</strong> any o<strong>the</strong>r part <strong>of</strong> <strong>the</strong> world.It is most strongly manifested in <strong>the</strong> Papilios and <strong>the</strong> Pierida?,and consists in tlie fore-wings being ei<strong>the</strong>r strongly curved orabruptly bent near <strong>the</strong> base, or in <strong>the</strong> extremity being elongatedand <strong>of</strong>ten somewhat hooked. Out <strong>of</strong> <strong>the</strong> 14 species <strong>of</strong> Papilioin Celebes, 13 exhibit this peculiarity in a greater or less degree,when compared with <strong>the</strong> most nearly allied species <strong>of</strong> <strong>the</strong>surrounding is<strong>land</strong>s. Ten species <strong>of</strong> Pieridse have <strong>the</strong> samecharacter, and in four or five <strong>of</strong> <strong>the</strong> Nymphalidse it is also verydistinctly marked. In almost every case <strong>the</strong> species found inCelebes are much larger than those <strong>of</strong> <strong>the</strong> is<strong>land</strong>s westward,and at least equal to those <strong>of</strong> <strong>the</strong> Moluccas, or even larger.<strong>The</strong> difierence <strong>of</strong> form is however <strong>the</strong> most remarkable feature,as it is altoge<strong>the</strong>r a new thing for a whole set <strong>of</strong> species in onecountry to clifier in exactly <strong>the</strong> same way from <strong>the</strong> correspondingsets in all <strong>the</strong> surrounding countries ; and it is so well marked,that without looking at <strong>the</strong> details <strong>of</strong> colouring, most CelebesPajDilios and many Pieridse can be at once distinguished fromthose <strong>of</strong> o<strong>the</strong>r is<strong>land</strong>s by <strong>the</strong>ir form alone.<strong>The</strong> outside figure <strong>of</strong> each pair here given shows <strong>the</strong> exactsize and form <strong>of</strong> <strong>the</strong> fore-wing in a buttei'fly <strong>of</strong> Celebes, while<strong>the</strong> inner one represents <strong>the</strong> most closely allied species fromone <strong>of</strong> <strong>the</strong> adjacent is<strong>land</strong>s. Figure 1 shows <strong>the</strong> stronglycurved margin <strong>of</strong> <strong>the</strong> Celebes species, Papilio gigon, comparedwith <strong>the</strong> much straighter margin <strong>of</strong> Papilio demolion fromSingapore and Java. Figure 2 shows <strong>the</strong> abruj^t bend over<strong>the</strong> base <strong>of</strong> <strong>the</strong> wing in Papilio miletus <strong>of</strong> Celebes comjDaredwith <strong>the</strong> slight curvature in <strong>the</strong> common Papilio sarjoedon,which has almost exactly <strong>the</strong> same form from India to NewGuinea and Australia. Figure 3 shows <strong>the</strong> elongated wing <strong>of</strong>Tachyris zarinda, a native <strong>of</strong> Celebes, compared with <strong>the</strong> muchshorter wing <strong>of</strong> Tachyris nero, a very closely allied species foundin all <strong>the</strong> western is<strong>land</strong>s. <strong>The</strong> diSerence <strong>of</strong> form is in each casesufficiently obvious, but when <strong>the</strong> insects <strong>the</strong>mselves are comparedit is much more striking than in <strong>the</strong>se partial outlines.From <strong>the</strong> analogy <strong>of</strong> birds, we should suppose that <strong>the</strong> pointedwing gave increased rapidity <strong>of</strong> flight, since it is a character <strong>of</strong>terns, swallows, falcons, and <strong>of</strong> <strong>the</strong> swift-flying pigeons. Ashort and rounded wing, on <strong>the</strong> o<strong>the</strong>r hand, always accompaniesa more feeble or more laborious flight, and one much less undercommand. We might suppose, <strong>the</strong>refore, that <strong>the</strong> butterflieswhich possess this peculiar form were better able to escape

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