The Malay archipelago : the land of the orang-utan ... - Wallace Online
The Malay archipelago : the land of the orang-utan ... - Wallace Online The Malay archipelago : the land of the orang-utan ... - Wallace Online
—214 THE MALAY ARCHIPELAGO. [chap.difference is as striking as anything can be. In the family ofthe Pieridte, or white butterflies, the difference is not quite sogreat, owing perhai^s to the more wandering habits of the group ;but it is still very remarkable. Out of 30 species inhabitingCelebes, 19 are peculiar, while Java (from which more speciesare known than from Sumatra or Borneo), out of 37 species hasonly 13 peculiar. The Danaidse are large, but weak-flyingbutterflies, ^vliich frequent forests and gardens, and are plainlybut often "\'ery richly coloured. Of these my own collectioncontains 16 species from Celebes and 15 from Borneo ; butwhereas no less than 14 are confined to the former island, onlytwo are peculiar to the latter. The Nymphalidse are a veryextensive group, of generally strong-winged and very brightcolouredbutterflies, very abundant in the tropics, and representedin our own country by our Fritillaries, our Vanessas,and our Purple-emperor. Some months ago I drew up a list ofthe Eastern species of this group, including all the new onesdiscovered by myself, and arrived at the following comparativeresults :species of Nyn^phaUd.. ^P^f^^lS'^^ of pfS^^/ecies.Java 70 23 33Borneo .... 52 15 . . • .... 29Celebes .... 48 35 73The ColeoiDtera are so extensive that few of the grouj^s haveyet been carefully worked out. I will therefore refer to oneonly, whicli I have myself recently studied—the Cetoniadfe orRose-chafers,—a group of beetles which, owing to their extremebeauty, have been much souglit after. From Java 37 sjDecies ofthese insects ai-e known, and from Celebes only 30 yet only13, or 35 per cent., are peculiar to the former island,;and 19, or63 per cent., to the latter.The result of these comparisons is, that although Celebes is asingle large island with only a few smaller ones closely groupedaround it, we must really consider it as forming one of thegreat divisions of the Arcliipelago, equal in i-ank and imjDoi'tanceto the whole of the ]\Ioluccan or Philippine groups, to thePajraan islands, or to the Indo-Malay islands (Java, Sumatra,Borneo, and the Malay peninsula). Taking those families ofinsects and birds which are best known, the following tableshows the comparison of Celebes with the other groups ofislands :PAPILIONID.E ANDHAWKS, PARROTS, ANDPIEBIDiE.PIGEONS.Percent, of peculiarPercent, of peculiarSpecies.Species.Iiido-Maliiy region 56 54Philippine group 66 73Celebes 69 60Moluccan group 52 62Timor group 42 47Papuan group 64 47
XVIII.] NATURAL HISTORY OF CELEBES. 215These large arid well-known families well represent thegeneral character of the zoology of Celebes ; and they showthat this island is really one of the most isolated joortions ofthe Archipelago, although situated in its very centre.But the insects of Celebes present us with other phenomenamore curious and more difficult to explain than their specificindividuality. The butterflies of that island ai'e in many casescharacterized by a peculiarity of outline, which distinguishesthem at a glance from those of any other part of the world.It is most strongly manifested in the Papilios and the Pierida?,and consists in tlie fore-wings being either strongly curved orabruptly bent near the base, or in the extremity being elongatedand often somewhat hooked. Out of the 14 species of Papilioin Celebes, 13 exhibit this peculiarity in a greater or less degree,when compared with the most nearly allied species of thesurrounding islands. Ten species of Pieridse have the samecharacter, and in four or five of the Nymphalidse it is also verydistinctly marked. In almost every case the species found inCelebes are much larger than those of the islands westward,and at least equal to those of the Moluccas, or even larger.The difierence of form is however the most remarkable feature,as it is altogether a new thing for a whole set of species in onecountry to clifier in exactly the same way from the correspondingsets in all the surrounding countries ; and it is so well marked,that without looking at the details of colouring, most CelebesPajDilios and many Pieridse can be at once distinguished fromthose of other islands by their form alone.The outside figure of each pair here given shows the exactsize and form of the fore-wing in a buttei'fly of Celebes, whilethe inner one represents the most closely allied species fromone of the adjacent islands. Figure 1 shows the stronglycurved margin of the Celebes species, Papilio gigon, comparedwith the much straighter margin of Papilio demolion fromSingapore and Java. Figure 2 shows the abruj^t bend overthe base of the wing in Papilio miletus of Celebes comjDaredwith the slight curvature in the common Papilio sarjoedon,which has almost exactly the same form from India to NewGuinea and Australia. Figure 3 shows the elongated wing ofTachyris zarinda, a native of Celebes, compared with the muchshorter wing of Tachyris nero, a very closely allied species foundin all the western islands. The diSerence of form is in each casesufficiently obvious, but when the insects themselves are comparedit is much more striking than in these partial outlines.From the analogy of birds, we should suppose that the pointedwing gave increased rapidity of flight, since it is a character ofterns, swallows, falcons, and of the swift-flying pigeons. Ashort and rounded wing, on the other hand, always accompaniesa more feeble or more laborious flight, and one much less undercommand. We might suppose, therefore, that the butterflieswhich possess this peculiar form were better able to escape
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—214 THE MALAY ARCHIPELAGO. [chap.difference is as striking as anything can be. In <strong>the</strong> family <strong>of</strong><strong>the</strong> Pieridte, or white butterflies, <strong>the</strong> difference is not quite sogreat, owing perhai^s to <strong>the</strong> more wandering habits <strong>of</strong> <strong>the</strong> group ;but it is still very remarkable. Out <strong>of</strong> 30 species inhabitingCelebes, 19 are peculiar, while Java (from which more speciesare known than from Sumatra or Borneo), out <strong>of</strong> 37 species hasonly 13 peculiar. <strong>The</strong> Danaidse are large, but weak-flyingbutterflies, ^vliich frequent forests and gardens, and are plainlybut <strong>of</strong>ten "\'ery richly coloured. Of <strong>the</strong>se my own collectioncontains 16 species from Celebes and 15 from Borneo ; butwhereas no less than 14 are confined to <strong>the</strong> former is<strong>land</strong>, onlytwo are peculiar to <strong>the</strong> latter. <strong>The</strong> Nymphalidse are a veryextensive group, <strong>of</strong> generally strong-winged and very brightcolouredbutterflies, very abundant in <strong>the</strong> tropics, and representedin our own country by our Fritillaries, our Vanessas,and our Purple-emperor. Some months ago I drew up a list <strong>of</strong><strong>the</strong> Eastern species <strong>of</strong> this group, including all <strong>the</strong> new onesdiscovered by myself, and arrived at <strong>the</strong> following comparativeresults :species <strong>of</strong> Nyn^phaUd.. ^P^f^^lS'^^ <strong>of</strong> pfS^^/ecies.Java 70 23 33Borneo .... 52 15 . . • .... 29Celebes .... 48 35 73<strong>The</strong> ColeoiDtera are so extensive that few <strong>of</strong> <strong>the</strong> grouj^s haveyet been carefully worked out. I will <strong>the</strong>refore refer to oneonly, whicli I have myself recently studied—<strong>the</strong> Cetoniadfe orRose-chafers,—a group <strong>of</strong> beetles which, owing to <strong>the</strong>ir extremebeauty, have been much souglit after. From Java 37 sjDecies <strong>of</strong><strong>the</strong>se insects ai-e known, and from Celebes only 30 yet only13, or 35 per cent., are peculiar to <strong>the</strong> former is<strong>land</strong>,;and 19, or63 per cent., to <strong>the</strong> latter.<strong>The</strong> result <strong>of</strong> <strong>the</strong>se comparisons is, that although Celebes is asingle large is<strong>land</strong> with only a few smaller ones closely groupedaround it, we must really consider it as forming one <strong>of</strong> <strong>the</strong>great divisions <strong>of</strong> <strong>the</strong> Arcliipelago, equal in i-ank and imjDoi'tanceto <strong>the</strong> whole <strong>of</strong> <strong>the</strong> ]\Ioluccan or Philippine groups, to <strong>the</strong>Pajraan is<strong>land</strong>s, or to <strong>the</strong> Indo-<strong>Malay</strong> is<strong>land</strong>s (Java, Sumatra,Borneo, and <strong>the</strong> <strong>Malay</strong> peninsula). Taking those families <strong>of</strong>insects and birds which are best known, <strong>the</strong> following tableshows <strong>the</strong> comparison <strong>of</strong> Celebes with <strong>the</strong> o<strong>the</strong>r groups <strong>of</strong>is<strong>land</strong>s :PAPILIONID.E ANDHAWKS, PARROTS, ANDPIEBIDiE.PIGEONS.Percent, <strong>of</strong> peculiarPercent, <strong>of</strong> peculiarSpecies.Species.Iiido-Maliiy region 56 54Philippine group 66 73Celebes 69 60Moluccan group 52 62Timor group 42 47Papuan group 64 47