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Introduction to Sports Biomechanics: Analysing Human Movement ...

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form muscle. The epimysium separates the muscle from its neighbours and facilitates<br />

frictionless movement.<br />

The contractile cells are concentrated in the soft, fleshy central part of the muscle,<br />

called the belly. Towards the ends of the muscle the contractile cells finish but the<br />

perimysium and epimysium continue <strong>to</strong> the bony attachment as a cord-like tendon or<br />

flat aponeurosis, in which the fibres are plaited <strong>to</strong> distribute the muscle force equally<br />

over the attachment area. If the belly continues almost <strong>to</strong> the bone, then individual<br />

sheaths of connective tissue attach <strong>to</strong> the bone over a larger area.<br />

The strength and thickness of the muscle sheath varies with location. Superficial<br />

muscles, particularly near the distal end of a limb have a thick sheath with additional<br />

protective fascia. The sheaths form a <strong>to</strong>ugh structural framework for the semi-fluid<br />

muscle tissue. They return <strong>to</strong> their original length even if stretched by 40% of their<br />

resting length. Groups of muscles are compartmentalised from others by intermuscular<br />

septa, usually attached <strong>to</strong> the bone and <strong>to</strong> the deep fascia that surrounds the muscles.<br />

Muscle activation<br />

Naming muscles<br />

THE ANATOMY OF HUMAN MOVEMENT<br />

Each muscle fibre is innervated by cranial or spinal nerves and is under voluntary<br />

control. The terminal branch of the nerve fibre ends at the neuromuscular junction or<br />

mo<strong>to</strong>r end-plate, which <strong>to</strong>uches the muscle fibre and transmits the nerve impulse <strong>to</strong> the<br />

sarcoplasm. Each muscle is entered from the central nervous system by nerves that<br />

contain both mo<strong>to</strong>r and sensory fibres, the former of which are known as mo<strong>to</strong>r<br />

neurons. As each mo<strong>to</strong>r neuron enters the muscle, it branches in<strong>to</strong> terminals, each of<br />

which forms a mo<strong>to</strong>r end-plate with a single muscle fibre. The term mo<strong>to</strong>r unit is used<br />

<strong>to</strong> refer <strong>to</strong> a mo<strong>to</strong>r neuron and all the muscle fibres that it innervates, and these can<br />

be spread over a wide area of the muscle. The mo<strong>to</strong>r unit can be considered the<br />

fundamental functional unit of neuromuscular control. Each nerve impulse causes all<br />

the muscle fibres of the mo<strong>to</strong>r unit <strong>to</strong> contract fully and almost simultaneously. The<br />

number of fibres per mo<strong>to</strong>r unit is sometimes called the innervation ratio. This ratio<br />

can be less than 10 for muscles requiring very fine control and greater than 1000 for the<br />

weight-bearing muscles of the lower extremity. Muscle activation is regulated through<br />

mo<strong>to</strong>r unit recruitment and the mo<strong>to</strong>r unit stimulation rate (or rate-coding). The<br />

former is an orderly sequence based on the size of the mo<strong>to</strong>r neuron. The smaller mo<strong>to</strong>r<br />

neurons are recruited first; these are typically slow twitch with a low maximum tension<br />

and a long contraction time. If more mo<strong>to</strong>r units can be recruited, then this mechanism<br />

dominates; smaller muscles have fewer mo<strong>to</strong>r units and depend more on increasing<br />

their stimulation rate.<br />

As noted in the introduction <strong>to</strong> this chapter, in the scientific literature muscles are<br />

nearly always known by their Latin names. The full name is musculus, which is often<br />

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