The geographical distribution of animals, with a study of the relations ...
The geographical distribution of animals, with a study of the relations ... The geographical distribution of animals, with a study of the relations ...
G4 DISTRIBUTION OF ANIMALS. [part i.the antipodes of each other, would be most inconvenient, evenif there were not such difference of opinion about them.Therecan be little doubt, for example, that the most radical zoologicaldivision of the earth is made by separating the Australian regionfrom the rest; but although it is something useful anddefinite to know that a group of animals is peculiar to Australia,it is exceedingly vague and unsatisfactory to say of any othergroup merely that it is extra-Australian. Neither can it be saidthat, from any point of view, these two divisions are of equalimportance. The next great natural division that can be madeis the separation of the Neotropical Region of Mr. Sclater fromthe rest of the world. We thus have three primary divisions,which Professor Huxley seems inclined to consider as oftolerably equal zoological importance. But a consideration ofall the facts, zoological and palseontological, indicates, that thegreat northern division (Arctogsea) is fully as much more importantthan either Australia or South America,as its four componentparts are less important ; and if so, convenience requiresus to adopt the smaller rather than the larger divisions.This question, of comparative importance or equivalence ofvalue, is very difficult to determine.It may be considered fromthe point of view of speciality or isolation, or from that ofrichness and variety of animal forms.In isolation and speciality,determined by what they want as well as what they possess, theAustralian and Neotropical regions are undoubtedly each comparablewith the rest of the earth (Arctogsea). But in richnessand variety of forms, they are both very much inferior, and aremuch more nearly comparable with the separate regions whichcompose it.Taking the families of mammalia as established bythe best authors, and leaving out the Cetacea and the Bats,which are almost universally distributed, and about whoseclassification there is much uncertainty, the number of familiesrepresented in each of Mr. Sclater's regions is as follows :I. Palsearctic region has 31 families of terrestrial mammalia.II. Ethiopian „ „ 40 „ „III. Indian „ „ 31 „ „IV. Australian „ „ 14 „ „V. Neotropical „ „ 26 „ „VI. Nearctic „ ,,23 „ „
;chap. iv.] ZOOLOGICAL REGIONS. 65We see, then, that even the exceedingly rich and isolated Neotropicalregion is less rich and diversified in its forms of mammalianlife than the very much smaller area of the Indian region,or the temperate Palcearctic, and very much less so than theEthiopian region ; while even the comparatively poor Nearcticregion, is nearly equal to it in the number of its family types. Ifthese were united they would possess fifty-five families, a numbervery disproportionate to those of the remaining two.Anotherconsideration is, that although the absence of certain forms oflife makes a region more isolated, it does not make itzoologicallymore important ; for we have only to suppose some five or sixfamilies, now common to both, to become extinct either in theEthiopian or the Indian regions, and they would become asstrongly differentiatedwhile still remaining as rich in family types.from all other regions as South America,In birds exactly thesame phenomenon recurs, the family types being less numerous inSouth America than in either of the other tropical regions of theearth, but a larger proportion of them are restricted to it.It willbe shown further on, that the Ethiopian and Indian, (or, as I proposeto call it in this work, Oriental) regions, are sufficiently differentiatedby very important groups of animals peculiar toeachand that, on strict zoological principles they are entitled torank asregions of equal value with the Neotropical and Australian.It is perhaps less clear whether the Palsearctic shouldbe separated from the Oriental region, with which it has undoubtedlymuch in common ; but there are many and powerfulreasons for keeping it distinct.facies in the animal forms of the two regions ;There is an unmistakably differentand although nofamilies of mammalia or birds, and not many genera, are whollyconfined to the Palsearctic region, a very considerable numberof both have their metropolis in it,and are very richly represented.The distinction between the characteristic forms of life in tropicaland cold countries is,on the whole, very strongly marked in thenorthern hemisphere ; and to refuse to recognise this in a subdivisionof the earth which is established for the very purpose ofexpressing such contrasts more clearly and concisely than byordinary geographicalterminology, would be both illogical and
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G4 DISTRIBUTION OF ANIMALS. [part i.<strong>the</strong> antipodes <strong>of</strong> each o<strong>the</strong>r, would be most inconvenient, evenif <strong>the</strong>re were not such difference <strong>of</strong> opinion about <strong>the</strong>m.<strong>The</strong>recan be little doubt, for example, that <strong>the</strong> most radical zoologicaldivision <strong>of</strong> <strong>the</strong> earth is made by separating <strong>the</strong> Australian regionfrom <strong>the</strong> rest; but although it is something useful anddefinite to know that a group <strong>of</strong> <strong>animals</strong> is peculiar to Australia,it is exceedingly vague and unsatisfactory to say <strong>of</strong> any o<strong>the</strong>rgroup merely that it is extra-Australian. Nei<strong>the</strong>r can it be saidthat, from any point <strong>of</strong> view, <strong>the</strong>se two divisions are <strong>of</strong> equalimportance. <strong>The</strong> next great natural division that can be madeis <strong>the</strong> separation <strong>of</strong> <strong>the</strong> Neotropical Region <strong>of</strong> Mr. Sclater from<strong>the</strong> rest <strong>of</strong> <strong>the</strong> world. We thus have three primary divisions,which Pr<strong>of</strong>essor Huxley seems inclined to consider as <strong>of</strong>tolerably equal zoological importance. But a consideration <strong>of</strong>all <strong>the</strong> facts, zoological and palseontological, indicates, that <strong>the</strong>great nor<strong>the</strong>rn division (Arctogsea) is fully as much more importantthan ei<strong>the</strong>r Australia or South America,as its four componentparts are less important ; and if so, convenience requiresus to adopt <strong>the</strong> smaller ra<strong>the</strong>r than <strong>the</strong> larger divisions.This question, <strong>of</strong> comparative importance or equivalence <strong>of</strong>value, is very difficult to determine.It may be considered from<strong>the</strong> point <strong>of</strong> view <strong>of</strong> speciality or isolation, or from that <strong>of</strong>richness and variety <strong>of</strong> animal forms.In isolation and speciality,determined by what <strong>the</strong>y want as well as what <strong>the</strong>y possess, <strong>the</strong>Australian and Neotropical regions are undoubtedly each comparable<strong>with</strong> <strong>the</strong> rest <strong>of</strong> <strong>the</strong> earth (Arctogsea). But in richnessand variety <strong>of</strong> forms, <strong>the</strong>y are both very much inferior, and aremuch more nearly comparable <strong>with</strong> <strong>the</strong> separate regions whichcompose it.Taking <strong>the</strong> families <strong>of</strong> mammalia as established by<strong>the</strong> best authors, and leaving out <strong>the</strong> Cetacea and <strong>the</strong> Bats,which are almost universally distributed, and about whoseclassification <strong>the</strong>re is much uncertainty, <strong>the</strong> number <strong>of</strong> familiesrepresented in each <strong>of</strong> Mr. Sclater's regions is as follows :I. Palsearctic region has 31 families <strong>of</strong> terrestrial mammalia.II. Ethiopian „ „ 40 „ „III. Indian „ „ 31 „ „IV. Australian „ „ 14 „ „V. Neotropical „ „ 26 „ „VI. Nearctic „ ,,23 „ „