The geographical distribution of animals, with a study of the relations ...

The geographical distribution of animals, with a study of the relations ... The geographical distribution of animals, with a study of the relations ...

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;G2 DISTRIBUTION OF ANIMALS. [part i.Mr. Sclater in his Lectures on Geographical Distribution at theZoological Gardens in May 1874), because it is absolutely withoutindigenous mammalia and very poor in all forms of life,and therefore by no means prominent or important enough toform a primary region of the earth.It may be as well here to notice what appears tobe a seriousobjection to making New Zealand, or any similar isolateddistrict, one of the great zoological regions, comparable to SouthAmerica, Australia, or Ethiopia ;which is, that its claim to thatdistinction rests on grounds which are liable to fail. It isbecause New Zealand, in addition to its negative merits, possessesthree families of birds (Apterygidse living, Dinornithidse andPalapterygidae extinct), and a peculiar lizard-like reptile,Hatteria, which has to be classed in a distinct order, Ehynchocephalina,that the rank of a Eegion is claimed for it. Butsupposing, what is not at all improbable, that other Ehynchocephalinashould be discovered in the interior of Australia orin New Guinea, and that Apterygidre or Palapterygi'dre shouldbe found to have inhabited Australia in Post-Pliocene times,(as Dinornithidae have already been proved to have done) theclaims of New Zealand would entirely fail, and it would beuniversally acknowledged to be a part of the great Australianregion. No such reversal can take place in the case of theother regions ;because they rest, not upon one or two, but upon alarge number of peculiarities, of such a nature that there isroom upon the globe for discoveries that can seriously modifythem. Even if one or two peculiar types, like Apterygidre ornoHatteria, should permanently remain characteristic of New Zealandalone, we can account for these by the extreme isolationofthe country, and the absence of enemies, which have enabledthese defenceless birds and reptiles to continue their existencejust as the isolation and protection of the caverns of Carniolahave enabled the Proteus to survive in Europe.But supposingthat the Proteus was the sole representative of an order ofBatrachia, and that two or three other equally curious andisolated forms occurred with it,no one would propose that thesecaverns or the district containing them, should form one of the

chap, iv.] ZOOLOGICAL REGIONS. 63primary divisions of the earth. Neither can much stress belaid on the negative characteristics of New Zealand, since theyare found to an almost equal extent in every oceanic island.Again, it is both inconvenient and misleading to pick outcertain tracts from the midst of one region or sub-region and toplace them in another, on account of certain isolated affinitieswhich may often be accounted for by local peculiarities.Evenif the resemblance of the fauna of Chili and Patagonia to thatof the Palsearctic and Nearctic regions was much greater than itis, this mode of dealing with it would be objectionable ; but itis still more so, when we find that these countries have astrongly marked South American character, and that the northernaffinities are altogether exceptional. The Eodentia, whichcomprise a large portion of the mammalia of these countries,are wholly South American in type, and the birds are almost allallied to forms characteristic of tropical America.For analogous reasons the Ethiopian must not be made toinclude any part of India or Ceylon ;for although the Fauna ofCentral India has some African affinities, these do not preponderate;and it will not be difficult to show that to follow Mr.Andrew Murray in uniting bodily the Ethiopian and Indianregions of Mr. Sclater, is both unnatural and inconvenient.Theresemblances between them are of the same character as thosewhich would unite them both with thePalasarctic and Nearcticregions ; and although it may be admitted, that, as ProfessorHuxley maintains, this group forms one of the great primarydivisions of the globe, it is far too extensive and too heterogeneousto subserve the practical uses for which we require adivision of the world into zoological regions.Reasons for adopting the six Regions first proposed by Mr. Sclater.—So that we do not violate any clear affinities or produce anyglaring irregularities, it is a positive, and by no means an unimportant,advantage to have our named regionsapproximatelyequal in size, and with easily defined, and therefore easily remembered,boundaries. All elaborate definitions of interpenetratingfrontiers, as well as regions extending over three-fourthsof the land surface of the globe, and including places which are

;G2 DISTRIBUTION OF ANIMALS. [part i.Mr. Sclater in his Lectures on Geographical Distribution at <strong>the</strong>Zoological Gardens in May 1874), because it is absolutely <strong>with</strong>outindigenous mammalia and very poor in all forms <strong>of</strong> life,and <strong>the</strong>refore by no means prominent or important enough t<strong>of</strong>orm a primary region <strong>of</strong> <strong>the</strong> earth.It may be as well here to notice what appears tobe a seriousobjection to making New Zealand, or any similar isolateddistrict, one <strong>of</strong> <strong>the</strong> great zoological regions, comparable to SouthAmerica, Australia, or Ethiopia ;which is, that its claim to thatdistinction rests on grounds which are liable to fail. It isbecause New Zealand, in addition to its negative merits, possessesthree families <strong>of</strong> birds (Apterygidse living, Dinornithidse andPalapterygidae extinct), and a peculiar lizard-like reptile,Hatteria, which has to be classed in a distinct order, Ehynchocephalina,that <strong>the</strong> rank <strong>of</strong> a Eegion is claimed for it. Butsupposing, what is not at all improbable, that o<strong>the</strong>r Ehynchocephalinashould be discovered in <strong>the</strong> interior <strong>of</strong> Australia orin New Guinea, and that Apterygidre or Palapterygi'dre shouldbe found to have inhabited Australia in Post-Pliocene times,(as Dinornithidae have already been proved to have done) <strong>the</strong>claims <strong>of</strong> New Zealand would entirely fail, and it would beuniversally acknowledged to be a part <strong>of</strong> <strong>the</strong> great Australianregion. No such reversal can take place in <strong>the</strong> case <strong>of</strong> <strong>the</strong>o<strong>the</strong>r regions ;because <strong>the</strong>y rest, not upon one or two, but upon alarge number <strong>of</strong> peculiarities, <strong>of</strong> such a nature that <strong>the</strong>re isroom upon <strong>the</strong> globe for discoveries that can seriously modify<strong>the</strong>m. Even if one or two peculiar types, like Apterygidre ornoHatteria, should permanently remain characteristic <strong>of</strong> New Zealandalone, we can account for <strong>the</strong>se by <strong>the</strong> extreme isolation<strong>of</strong><strong>the</strong> country, and <strong>the</strong> absence <strong>of</strong> enemies, which have enabled<strong>the</strong>se defenceless birds and reptiles to continue <strong>the</strong>ir existencejust as <strong>the</strong> isolation and protection <strong>of</strong> <strong>the</strong> caverns <strong>of</strong> Carniolahave enabled <strong>the</strong> Proteus to survive in Europe.But supposingthat <strong>the</strong> Proteus was <strong>the</strong> sole representative <strong>of</strong> an order <strong>of</strong>Batrachia, and that two or three o<strong>the</strong>r equally curious andisolated forms occurred <strong>with</strong> it,no one would propose that <strong>the</strong>secaverns or <strong>the</strong> district containing <strong>the</strong>m, should form one <strong>of</strong> <strong>the</strong>

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