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The geographical distribution of animals, with a study of the relations ...

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54 DISTRIBUTION OF ANIMALS. [part i.and this, it may be said, proves <strong>the</strong>ir fundamental unity andthat <strong>the</strong>y ought to form but one primary region. To obviatesome <strong>of</strong> <strong>the</strong>se difficulties a binary or dichotomous division issometimes proposed; that portion <strong>of</strong> <strong>the</strong> earth which differsmost from <strong>the</strong> rest being cut <strong>of</strong>f as a region equal in rankto all that remains, which is subjected again and again to<strong>the</strong> same process.To decide <strong>the</strong>se various points it seems advisable that convenience,intelligibility, and custom, should largely guide us.<strong>The</strong> first essential is, a broadly marked and easily rememberedset <strong>of</strong> regions ;which correspond, as nearly as truth to naturewill allow, <strong>with</strong> <strong>the</strong> <strong>distribution</strong> <strong>of</strong> <strong>the</strong> most important groups<strong>of</strong> <strong>animals</strong>.What <strong>the</strong>se groups are we shall presently explain.In determining <strong>the</strong> number, extent, and boundaries <strong>of</strong> <strong>the</strong>seregions, we must be guided by a variety <strong>of</strong> indications, since<strong>the</strong> application <strong>of</strong> fixed rules is impossible. <strong>The</strong>y should evidentlybe <strong>of</strong> a moderate number, corresponding as far aspracticable <strong>with</strong> <strong>the</strong> great natural divisions <strong>of</strong> <strong>the</strong> globe markedout by nature, and which have always been recognized bygeographers. <strong>The</strong>re should be some approximation to equality<strong>of</strong> size, since <strong>the</strong>re is reason to believe that a tolerably extensivearea has been an essential condition for <strong>the</strong> development <strong>of</strong>most animal forms ; and it is found that, o<strong>the</strong>r things beingequal, <strong>the</strong> numbers, variety and importance <strong>of</strong> <strong>the</strong> forms <strong>of</strong>animal and vegetable life, do bear some approximate relationto extent <strong>of</strong> area. Although <strong>the</strong> possession <strong>of</strong> peculiar familiesor genera is<strong>the</strong> main character <strong>of</strong> a primary zoological region,yet <strong>the</strong> negative character <strong>of</strong> <strong>the</strong> absence <strong>of</strong> certain familiesor genera is <strong>of</strong> equal importance, when this absence does notmanifestly depend on unsuitability to <strong>the</strong> support <strong>of</strong> <strong>the</strong> group,and especially when <strong>the</strong>re is now no physical barrier preventing<strong>the</strong>ir entrance.This will become evident when we consider that<strong>the</strong> importance <strong>of</strong> <strong>the</strong> possession <strong>of</strong> a group by one region dependson its absence from <strong>the</strong> adjoining regions ; and if <strong>the</strong>re isnow no barrier to its entrance, we may be sure that <strong>the</strong>re hasonce been one ;and that <strong>the</strong> possession <strong>of</strong> <strong>the</strong> area by a distinctand well balanced set <strong>of</strong> organisms, which must have been slowly

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