The geographical distribution of animals, with a study of the relations ...

The geographical distribution of animals, with a study of the relations ... The geographical distribution of animals, with a study of the relations ...

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42 DISTRIBUTION OF ANIMALS. [part iamong forms implying a climate very little different from thepresent ; and our own Crag formation furnishes evidence of agradual refrigeration of climate ; since its three divisions, theCoralline, Ked, and Norwich Crags, show a decreasing numberof southern, and an increasing number of northern species, as weapproach the Glacial epoch. Still later than these we have theshells of the drift, almost all of which are northern and manyof them arctic species. Among the mammalia indicative ofcold, are the mammoth and the reindeer.In gravels and cavedepositsof Post-Pliocene date we find the same two animals,which soon disappear asthe climate approached its present condition;and Professor Forbes has given a list of fifty shellswhich inhabited the British seas before the Glacial epoch andinhabit it still, but are all wanting in the glacial deposits. Thewhole of these are found in the Newer Pliocene strata of Sicilyand the south of Europe, where they escaped destruction duringthe glacial winter.There are also certain facts in the distribution of plants, whichare so well explained by the Glacial epoch that they may be saidto give an additional confirmation to it. All over the northernhemisphere within the glaciated districts, the summits of loftymountains produce plants identical with those of the polarregions. In the celebrated case of the "White Mountains an NewHampshire, United States (latitude 45°), all the plants on thesummit are arctic species, none of which exist in the lowlandsfor near a thousand miles further north. It has also been remarked.that the plants of each mountain are more especiallyrelated to those of the countries directly north of it. Thus,those of the Pyrenees and of Scotland are Scandinavian, andthose of the White Mountains are all species found in Labrador.Now, remembering that we have evidence of an exceedinglymild and uniform climate in the arctic regions during theMiocene period and aevident that with each degreegradual refrigeration from that time, it isof change more and more hardyplants would be successively driven southwards ;till at last theplains of the temperate zone would be inhabited by plants, whichwere once confined to alpine heights or to the arctic regions.

chap, ill.] CONDITIONS AFFECTING DISTRIBUTION. 43As the icy mantle gradually melted off the face of the earththese plants would occupy the newly exposed soil, and wouldthus necessarily travel in two directions, back towards the arcticcircle and up towards the alpine peaks. The facts are thusexactly explained by a cause which independent evidence hasproved to be a real one, and every such explanation is an additionalproof of the reality of the cause. But this explanation implies,that in cases where the Glacial epoch cannot have so actedalpine plants should not be northern plants ;and a striking proofof this is to be found on the Peak of Teneriffe, a mountain12,000 feet high. In the uppermost 4,500 feet of this mountainabove the limit of trees, Von Buch found only eleven species ofplants, eight of which were peculiar ; but the whole were alliedto those found at lower elevations. On the Alps or Pyrenees atthis elevation, there would be a rich flora comprising hundredsof arctic plants ; and the absence of anything corresponding tothem in this case, in which their ingress was cut off by the sea,is exactly what the theory leads us to expect.Changes of Vegetation as affecting the Distribution of Animals.—As so many animals are dependent on vegetation, its changesimmediately affect their distribution. A remarkable example ofthis is afforded by the pre-historic condition of Denmark, asinterpreted by means of the peat-bogs and kitchen-middens.This country is now celebrated for itsbeech-trees; oaks and pinesbeing scarce ; and it is known to have had the same vegetation inthe time of the Eomans. In the peat-bogs, however, are founddeposits of oak trees ; and deeper still pines alone occur. Nowthe kitchen-middens tell us much of the natural history ofDenmark in the early Stone period ; and a curious confirmationof the fact that Denmark like Norway was then chiefly coveredwith pine forests is obtained by the discovery, that the Capercailziewas then abundant, a bird which feeds almost exclusivelyon the young shoots and seeds of pines and allied plants. Thecause of this change in the vegetation is unknown ; but from theknown fact that when forests are destroyed trees, of a differentkind usually occupy the ground, we may suppose that some suchchange as a temporary submergence might cause an entirely

chap, ill.] CONDITIONS AFFECTING DISTRIBUTION. 43As <strong>the</strong> icy mantle gradually melted <strong>of</strong>f <strong>the</strong> face <strong>of</strong> <strong>the</strong> earth<strong>the</strong>se plants would occupy <strong>the</strong> newly exposed soil, and wouldthus necessarily travel in two directions, back towards <strong>the</strong> arcticcircle and up towards <strong>the</strong> alpine peaks. <strong>The</strong> facts are thusexactly explained by a cause which independent evidence hasproved to be a real one, and every such explanation is an additionalpro<strong>of</strong> <strong>of</strong> <strong>the</strong> reality <strong>of</strong> <strong>the</strong> cause. But this explanation implies,that in cases where <strong>the</strong> Glacial epoch cannot have so actedalpine plants should not be nor<strong>the</strong>rn plants ;and a striking pro<strong>of</strong><strong>of</strong> this is to be found on <strong>the</strong> Peak <strong>of</strong> Teneriffe, a mountain12,000 feet high. In <strong>the</strong> uppermost 4,500 feet <strong>of</strong> this mountainabove <strong>the</strong> limit <strong>of</strong> trees, Von Buch found only eleven species <strong>of</strong>plants, eight <strong>of</strong> which were peculiar ; but <strong>the</strong> whole were alliedto those found at lower elevations. On <strong>the</strong> Alps or Pyrenees atthis elevation, <strong>the</strong>re would be a rich flora comprising hundreds<strong>of</strong> arctic plants ; and <strong>the</strong> absence <strong>of</strong> anything corresponding to<strong>the</strong>m in this case, in which <strong>the</strong>ir ingress was cut <strong>of</strong>f by <strong>the</strong> sea,is exactly what <strong>the</strong> <strong>the</strong>ory leads us to expect.Changes <strong>of</strong> Vegetation as affecting <strong>the</strong> Distribution <strong>of</strong> Animals.—As so many <strong>animals</strong> are dependent on vegetation, its changesimmediately affect <strong>the</strong>ir <strong>distribution</strong>. A remarkable example <strong>of</strong>this is afforded by <strong>the</strong> pre-historic condition <strong>of</strong> Denmark, asinterpreted by means <strong>of</strong> <strong>the</strong> peat-bogs and kitchen-middens.This country is now celebrated for itsbeech-trees; oaks and pinesbeing scarce ; and it is known to have had <strong>the</strong> same vegetation in<strong>the</strong> time <strong>of</strong> <strong>the</strong> Eomans. In <strong>the</strong> peat-bogs, however, are founddeposits <strong>of</strong> oak trees ; and deeper still pines alone occur. Now<strong>the</strong> kitchen-middens tell us much <strong>of</strong> <strong>the</strong> natural history <strong>of</strong>Denmark in <strong>the</strong> early Stone period ; and a curious confirmation<strong>of</strong> <strong>the</strong> fact that Denmark like Norway was <strong>the</strong>n chiefly covered<strong>with</strong> pine forests is obtained by <strong>the</strong> discovery, that <strong>the</strong> Capercailziewas <strong>the</strong>n abundant, a bird which feeds almost exclusivelyon <strong>the</strong> young shoots and seeds <strong>of</strong> pines and allied plants. <strong>The</strong>cause <strong>of</strong> this change in <strong>the</strong> vegetation is unknown ; but from <strong>the</strong>known fact that when forests are destroyed trees, <strong>of</strong> a differentkind usually occupy <strong>the</strong> ground, we may suppose that some suchchange as a temporary submergence might cause an entirely

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