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The geographical distribution of animals, with a study of the relations ...

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chap, xiii.] THE AUSTRALIAN REGION. 437finding <strong>the</strong> country already fairly stocked, comparatively fewgroups were able to establish <strong>the</strong>mselves.Going back a little far<strong>the</strong>r, we come to <strong>the</strong> entrance <strong>of</strong> thosefew birds and insects which belong to India or Indo-China ;andthis probably occurred at <strong>the</strong> same time as that continentalextension southward, which we found was required to account fora similar phenomenon in Java. Celebes, being more remote,received only a few stragglers. We have now to go muchfar<strong>the</strong>r back, to <strong>the</strong> time when <strong>the</strong> ancestors <strong>of</strong> <strong>the</strong> peculiarCelebesian genera entered <strong>the</strong> country, and here our conjecturesmust necessarily be less defined.On <strong>the</strong> Australian side we have to account for Megacephalon,and <strong>the</strong> o<strong>the</strong>r genera <strong>of</strong> purely Papuan type. It may perhapsbe sufficient to say, that we do not yet know that <strong>the</strong>se genera,or some very close allies, do not still exist in New Guinea ; inwhich case <strong>the</strong>y may well have entered at <strong>the</strong> same time <strong>with</strong><strong>the</strong> species, already referred to. If, on <strong>the</strong> o<strong>the</strong>r hand, <strong>the</strong>y arereally as isolated as <strong>the</strong>y appear to be, <strong>the</strong>y representan earliercommunication, ei<strong>the</strong>r by an approximation <strong>of</strong> <strong>the</strong> two islandsover <strong>the</strong> space now occupied by <strong>the</strong> Moluccas ; or, what is perhapsmore probable, through a former extension <strong>of</strong> <strong>the</strong> Moluccas,which have since undergone so much subsidence, as to lead to<strong>the</strong> extinction <strong>of</strong> a large proportion <strong>of</strong> <strong>the</strong>ir ancient fauna.<strong>The</strong> wide-spread volcanic action, and especially <strong>the</strong> prevalence <strong>of</strong>raised coral-reefs in almost all <strong>the</strong> islands, render this lastsupposition very probable.On <strong>the</strong> Oriental side <strong>the</strong> difficulty is greater ; for here we find,what seem to be clear indications <strong>of</strong> a connection <strong>with</strong> Africa, aswell as <strong>with</strong> Continental Asia, at some immensely remote epoch.Cynopi<strong>the</strong>cus, Babirusa, and Anoa; Ceijcopsis, Streptocitta, and Gazzola(s. g.), and perhaps Scissirostrum, may be well explained asdescendants <strong>of</strong> ancestral types in<strong>the</strong>ir respective groups, whichalso gave rise to <strong>the</strong> special forms <strong>of</strong> Africa on <strong>the</strong> one hand, and <strong>of</strong>Asia on <strong>the</strong> o<strong>the</strong>r.For this immigration we must suppose, that ata period before <strong>the</strong> formation <strong>of</strong> <strong>the</strong> present Indo-Malay Islands,a great tract <strong>of</strong> land extended in anorth-westerly direction, tillit met <strong>the</strong> old Asiatic continent. This may have been before

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