The geographical distribution of animals, with a study of the relations ...

The geographical distribution of animals, with a study of the relations ... The geographical distribution of animals, with a study of the relations ...

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:424 ZOOLOGICAL GEOGRAPHY. [part hi.Moluccas (with New Guinea and Celebes,) Australia, and Java.The Moluccan forms may well have arrived as stragglers fromisland to island, aided by whatever facilitiesby lands now submerged.have been affordedMost of the remainder have been derivedeither from Australia or from Java ; and as their relationsto these islands are very interesting, they must be discussedwith some detail.Origin of the Timorese Fauna. — We must first note, that 80species, or exactly one-half of the land-birdsof the islands, arepeculiar and mostly very distinct, intimating that the immigrationcommenced long enough back to allow of much specificmodification. There is also one peculiar genus of kingfishers,Cariclonax, found only in Lombok and Flores, and more alliedto Australian than to Oriental types. The fine white-bandedpigeons (s. g. Leucotreron) are also almost peculiar ;one otherless typical species only being known, a native of N. Celebes.In order to compare thespecies with regard to their origin, wemust first take away those of wide distributionspecial indications can be obtained.from which noIn this case 49 of the landbirdsmust be deducted, leaving 111 species which affordgoodmaterials for comparison. These, when traced to their origin,show that 62 came from some part of the Australian region, 49from Java or the Oriental region. But if we divide them intotwo groups, the one containing the species identical with thoseof the Australian or Oriental regions, the other containing alliedor representative species peculiar to the islands, we have the followingresultSpecies common to the Timorese Islands and the Oriental Region 30Peculiar Timorese species allied to those of the. Oriental Region 19Total 49Species common to the Timorese Islands and the AustralianEegion 18Peculiar Timorese species allied to those of the Australian Region 44Total , 62This table is very important, as indicating that the connection

chap, xiii.] THE AUSTRALIAN REGION. 425with Australia was probably earlier than that with Java ; sincethe majority of the Australian species have become modified,while the majority of the Oriental species have remained unchanged.This is due, no doubt, in part to the continued immigrationof fresh individuals from Java, after that from Australia,the Moluccas and New Guinea had almost wholly ceased.must also notice the very small proportion of the genera,Weeitherof Australia or Java, that have found their way into these islands,many of the largest and most wide-spread groups in both countriesbeing altogether absent.Taking these facts into consideration,it is pretty clear that there has been no close and longcontinuedapproximation of these islands to any part of theAustralian region ; and it is also probable that they were fairlystocked with such Australian groups as they possess before theimmigration from Java commenced, or a largernumber of characteristicOriental forms would have been able to have establishedthemselves.On looking at our map, we find that a shallow submerged bankextends from Australia to within about twenty miles of the coastof Timor; and this is probably an indication that the twocountries were once only so far apart.This would have allowedthe purely Australian types to enter, as they are not numerous;there being about 6 Australian species,and 10 or 12 representativesof Australian species, in Timor.All the rest may have beenderived from the Moluccas or New Guinea, being mostly widespreadgenera of the Australian region; and the extension ofPapua in a south-west direction towards Java (which was suggestedas a means of providing New Guinea with peculiar Indo-Malay types not found in any other part of the region) mayhave probably served to supply Timor and Flores with the massof theirthe sea.Austro-Malayan genera across a narrow strait or arm ofLombok, Baly, and Sumbawa were probably not thenin existence, or nothing more than small \olcanic cones risingout of the sea, thus leaving a distance of 300 miles betweenFlores and Java. Subsequently they grew into islands, whichoffered an easy passage for a number of Indo-Malay generainto such scantily stocked territories as Flores and Timor. TheVol. I.—29

:424 ZOOLOGICAL GEOGRAPHY. [part hi.Moluccas (<strong>with</strong> New Guinea and Celebes,) Australia, and Java.<strong>The</strong> Moluccan forms may well have arrived as stragglers fromisland to island, aided by whatever facilitiesby lands now submerged.have been affordedMost <strong>of</strong> <strong>the</strong> remainder have been derivedei<strong>the</strong>r from Australia or from Java ; and as <strong>the</strong>ir <strong>relations</strong>to <strong>the</strong>se islands are very interesting, <strong>the</strong>y must be discussed<strong>with</strong> some detail.Origin <strong>of</strong> <strong>the</strong> Timorese Fauna. — We must first note, that 80species, or exactly one-half <strong>of</strong> <strong>the</strong> land-birds<strong>of</strong> <strong>the</strong> islands, arepeculiar and mostly very distinct, intimating that <strong>the</strong> immigrationcommenced long enough back to allow <strong>of</strong> much specificmodification. <strong>The</strong>re is also one peculiar genus <strong>of</strong> kingfishers,Cariclonax, found only in Lombok and Flores, and more alliedto Australian than to Oriental types. <strong>The</strong> fine white-bandedpigeons (s. g. Leucotreron) are also almost peculiar ;one o<strong>the</strong>rless typical species only being known, a native <strong>of</strong> N. Celebes.In order to compare <strong>the</strong>species <strong>with</strong> regard to <strong>the</strong>ir origin, wemust first take away those <strong>of</strong> wide <strong>distribution</strong>special indications can be obtained.from which noIn this case 49 <strong>of</strong> <strong>the</strong> landbirdsmust be deducted, leaving 111 species which affordgoodmaterials for comparison. <strong>The</strong>se, when traced to <strong>the</strong>ir origin,show that 62 came from some part <strong>of</strong> <strong>the</strong> Australian region, 49from Java or <strong>the</strong> Oriental region. But if we divide <strong>the</strong>m intotwo groups, <strong>the</strong> one containing <strong>the</strong> species identical <strong>with</strong> those<strong>of</strong> <strong>the</strong> Australian or Oriental regions, <strong>the</strong> o<strong>the</strong>r containing alliedor representative species peculiar to <strong>the</strong> islands, we have <strong>the</strong> followingresultSpecies common to <strong>the</strong> Timorese Islands and <strong>the</strong> Oriental Region 30Peculiar Timorese species allied to those <strong>of</strong> <strong>the</strong>. Oriental Region 19Total 49Species common to <strong>the</strong> Timorese Islands and <strong>the</strong> AustralianEegion 18Peculiar Timorese species allied to those <strong>of</strong> <strong>the</strong> Australian Region 44Total , 62This table is very important, as indicating that <strong>the</strong> connection

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