The geographical distribution of animals, with a study of the relations ...
The geographical distribution of animals, with a study of the relations ... The geographical distribution of animals, with a study of the relations ...
420 ZOOLOGICAL GEOGRAPHY. [part hi.Taking all the facts into consideration, I would suggest as themost probable explanation, that if the Moluccas ever formed partof the main Papuan land, they were separated at an early date,and subsequently so greatly submerged as to destroy a largeproportion of their fauna. They have since risen, and haveprobably been larger than at present, and rather more closelyapproximated to the parent land, whence they received a considerableimmigration of such animals as were adapted to crossnarrow seas. This gave them several Papuan forms, but stillleft them without a number of the types more especially confinedto the forest depths, or powerful enough to combat thegales which often blow weaker flyers out to sea. Most of thebirds whose absence from the Moluccas is so conspicuous belongto one or other of these classes.Among the most characteristic birds of the Moluccas are thehandsome crimson lories of the genera Lorius and Eos. Theseare found in every island (but not in Celebes or the Timorgroup) ;and a fine species of Eos, peculiar to the small islands ofSiau and Sanguir, just north of Celebes, obliges us to placethese with the Moluccas instead of with the former island, towhich they seem most naturally to belong.The crimson parrotsof the genus Eclectus are almost equally characteristic of theMoluccas, and add greatly to the brilliancy of the ornithology ofthese favoured islands.Reptiles.—The Reptiles, so far as known, appear to agree intheir distribution with the other vertebrates. In some smallcollections from Ceram there were no less than six of the generapeculiar to the Australian region, and which were before onlyknofn from Australia itself. These are, of snakes, Liasis andEnygrus, genera of Pythonidae ;with Diemenia and Acanthophis(Elapidae) ; of lizards, Cyclodus, a genus of Scincidae ; and ofAmphibia, a tree-frog of the genus Pelodryas.Insects— Peculiarities of the Moluccan Fauna.—In insects theMoluccas are hardly, if at all, inferior to New Guinea itself.Theislands abound in grand Papilios of the largest size and extremebeauty ;and it is a very remarkable fact, that when the closelyalliedspecies of the Moluccas and New Guinea arecompared,
;chap. xiii. J• THEAUSTRALIAN REGION. 421the former are almost always the largest.As examples may bementioned, Ornithoptera priamus and 0. helena of the Moluccas,both larger than the varieties (or species) of Papua ; Papilioulysses and deiplwbus of Amboyna, usually larger than theirallies in New Guinea ; Hestia idea, the largest species of thegenus ; Diadema pandarus and Charaxes euryalus, both largerthan any other species of the same genera in the whole archipelago.It is to be noted also, that in the Moluccas, the verylargest specimens or races seem always toisland of Amboyna ;come from the smalleven those of Ceram, the much larger islandto which it is a satellite, being almost always of less dimensions.Among Coleoptera, the Moluccas produce Euchirus longimanus,one of the largest and most remarkable of the LamellicornesSphingnotus dunningi, the largest of the Austro-Malayan Tmesisterninas;a Sphemira, the largest and handsomest of an extensivegenus ; an unusually large Schizorhina (Cetoniidse) ; andsome of the most remarkable and longest-horned Anthotribidoe.Even in birds the same law may be seen at work,—in the Tanysipteranais of Ceram, which has a larger tailthan any other inthe genus ;in Centropus goliath of Gilolo, being the largest andlongest-tailed species ; in Hydrornis maximus of Gilolo, thelargest and perhaps the most elegantly and conspicuouslycoloured of all the Pittidse ; in Platycercus amboinensis, beingpre-eminent in its ample blue tail ; in the two Moluccan loriesand Eos rubra, being more conspicuously red than the allied NewGuinea species ;and in Mcgapodius wallacei of Bouru, being theonly species of the genus conspicuously marked and banded.All these examples, of larger size, of longer tails or otherappendages, and of more conspicuous colouring, are probablyindications of a less severe struggle for existence in these islandsthan in the larger tract of New Guinea, with a more abundantand more varied fauna ; and this may apply even to the smallerislands, as compared with the larger in the immediate vicinity.The limited number of forms in the small islands comparedwith a similar area in the parent land, implies, perhaps, lesscompetition and less danger ; and thus allows, where all otherconditions are favourable, an unchecked and continuous de-
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420 ZOOLOGICAL GEOGRAPHY. [part hi.Taking all <strong>the</strong> facts into consideration, I would suggest as <strong>the</strong>most probable explanation, that if <strong>the</strong> Moluccas ever formed part<strong>of</strong> <strong>the</strong> main Papuan land, <strong>the</strong>y were separated at an early date,and subsequently so greatly submerged as to destroy a largeproportion <strong>of</strong> <strong>the</strong>ir fauna. <strong>The</strong>y have since risen, and haveprobably been larger than at present, and ra<strong>the</strong>r more closelyapproximated to <strong>the</strong> parent land, whence <strong>the</strong>y received a considerableimmigration <strong>of</strong> such <strong>animals</strong> as were adapted to crossnarrow seas. This gave <strong>the</strong>m several Papuan forms, but stillleft <strong>the</strong>m <strong>with</strong>out a number <strong>of</strong> <strong>the</strong> types more especially confinedto <strong>the</strong> forest depths, or powerful enough to combat <strong>the</strong>gales which <strong>of</strong>ten blow weaker flyers out to sea. Most <strong>of</strong> <strong>the</strong>birds whose absence from <strong>the</strong> Moluccas is so conspicuous belongto one or o<strong>the</strong>r <strong>of</strong> <strong>the</strong>se classes.Among <strong>the</strong> most characteristic birds <strong>of</strong> <strong>the</strong> Moluccas are <strong>the</strong>handsome crimson lories <strong>of</strong> <strong>the</strong> genera Lorius and Eos. <strong>The</strong>seare found in every island (but not in Celebes or <strong>the</strong> Timorgroup) ;and a fine species <strong>of</strong> Eos, peculiar to <strong>the</strong> small islands <strong>of</strong>Siau and Sanguir, just north <strong>of</strong> Celebes, obliges us to place<strong>the</strong>se <strong>with</strong> <strong>the</strong> Moluccas instead <strong>of</strong> <strong>with</strong> <strong>the</strong> former island, towhich <strong>the</strong>y seem most naturally to belong.<strong>The</strong> crimson parrots<strong>of</strong> <strong>the</strong> genus Eclectus are almost equally characteristic <strong>of</strong> <strong>the</strong>Moluccas, and add greatly to <strong>the</strong> brilliancy <strong>of</strong> <strong>the</strong> ornithology <strong>of</strong><strong>the</strong>se favoured islands.Reptiles.—<strong>The</strong> Reptiles, so far as known, appear to agree in<strong>the</strong>ir <strong>distribution</strong> <strong>with</strong> <strong>the</strong> o<strong>the</strong>r vertebrates. In some smallcollections from Ceram <strong>the</strong>re were no less than six <strong>of</strong> <strong>the</strong> generapeculiar to <strong>the</strong> Australian region, and which were before onlykn<strong>of</strong>n from Australia itself. <strong>The</strong>se are, <strong>of</strong> snakes, Liasis andEnygrus, genera <strong>of</strong> Pythonidae ;<strong>with</strong> Diemenia and Acanthophis(Elapidae) ; <strong>of</strong> lizards, Cyclodus, a genus <strong>of</strong> Scincidae ; and <strong>of</strong>Amphibia, a tree-frog <strong>of</strong> <strong>the</strong> genus Pelodryas.Insects— Peculiarities <strong>of</strong> <strong>the</strong> Moluccan Fauna.—In insects <strong>the</strong>Moluccas are hardly, if at all, inferior to New Guinea itself.<strong>The</strong>islands abound in grand Papilios <strong>of</strong> <strong>the</strong> largest size and extremebeauty ;and it is a very remarkable fact, that when <strong>the</strong> closelyalliedspecies <strong>of</strong> <strong>the</strong> Moluccas and New Guinea arecompared,