The geographical distribution of animals, with a study of the relations ...
The geographical distribution of animals, with a study of the relations ... The geographical distribution of animals, with a study of the relations ...
402 ZOOLOGICAL GEOGRAPHY. [part hi.not only on account of what we know of the permanence ofcontinents and deep oceans, hut because such a connection musthave led to much more numerous and important casesof similarityof natural productions than we actually find. And ifwithin the life of species such interchange may have takenplace across seas of greater or less extent, still more easy is itto understand, how, within the life of genera and families, a numberof such interchanges may have occurred ;yet always limitedto those groups whose conditions of life render transmissionpossible. Had an actual land connection existed within thetemperate zone, or during a period of warmth in the Antarcticregions, there would have been no such strict limitations to theinter-migration of animals.that floating ice has had some share in theIt may be held to support the viewtransmission of fishand amphibia, when -we find that in the case of the narrowtropical sea dividing Borneo from Celebes and the Moluccas, noproportionate amount of transmission has taken place, butnumerous species, genera, and whole families, terminate abruptlyat what we have other reasons for believing to be the furthestlimits of an ancient continent. We can hardly suppose, however,that this mode of transmission would have sufficed forsuch groups as tree-frogs, which are inhabitants of the moretemperate or even warm portions of the two southern lands.Some of these cases may perhaps be explained by the suppositionof a considerable extent of land in the South-Temperate andAntarctic regions now submerged, and by a warm or temperateclimate analogous to that which prevailed in the Arctic regionsduring some part of the Miocene epoch ; while others may bedue to cases of survival in the two areas of once wide-spreadgroups, a view supported in the case of the Amphibia by theerratic manner in which many of the groups are spread overthe globe.From an examination of the facts presented by the variousclasses of vertebrates, we are, then, led to the conclusion,that there is no evidence of a former land-connection be,tween the Australian and Neotropical regions ;but that thevarious scattered resemblances in their natural productions
chap, xiii.] THE AUSTKALIAN REGION. 403that undoubtedly occur, are probably due to three distinctcauses.First, we have the American Didelphyida?, among Mammals,and the Cracidse, among birds, allied respectively to the Marsupialsand the Megapodiidsp, of Australia. This is probably morea coincidence than an affinity,due to the preservation of ancientwide-spread types in two remote areas, each cut off from thegreat northern continental masses, inwhich higher forms wereevolved leading to the extinction of the lower types. In eachof these southern isolated lands the original type would undergoa special development ; in the one case suited to an arborealexistence, in the other to a life among arid plains.The second case is that of the tree-frogs, and the genus Osteorjlossumamong fishes ; and is most likely due to the extensionand approximation of the two southern continents, and the existenceof some intermediate lands, during a warm period whenfacilities would be afforded for the transmission of a few organismsby the causes which have led to the exceptional diffusionof fresh-water productions in all parts of the world. As howeverOsteoglossum occurs also in the Sunda Islands, this may be a caseof survival of a once wide-spread group.The third case is that of the same genera and even species offish, and perhaps of frogs, in the two countries ; which maybe due to transmission from island to island by the aid of floatingice, with or without the assistance of more interveninglands than now exist.Having arrived at these conclusions from a consideration ofthe vertebrata, we shall be in a position to examine how farthe same causes will explain, or agree with, the distribution of theinvertebrate groups, or elucidate any special difficulties we maymeet with in the relations of the sub-regions.Insects.The insects of the Australian region are as varied, and insome respects as peculiar as its higher forms of life. As wehave already indicated in our sketch of the Oriental region, avast number of forms inhabit the Austro-Malay sub-region
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402 ZOOLOGICAL GEOGRAPHY. [part hi.not only on account <strong>of</strong> what we know <strong>of</strong> <strong>the</strong> permanence <strong>of</strong>continents and deep oceans, hut because such a connection musthave led to much more numerous and important cases<strong>of</strong> similarity<strong>of</strong> natural productions than we actually find. And if<strong>with</strong>in <strong>the</strong> life <strong>of</strong> species such interchange may have takenplace across seas <strong>of</strong> greater or less extent, still more easy is itto understand, how, <strong>with</strong>in <strong>the</strong> life <strong>of</strong> genera and families, a number<strong>of</strong> such interchanges may have occurred ;yet always limitedto those groups whose conditions <strong>of</strong> life render transmissionpossible. Had an actual land connection existed <strong>with</strong>in <strong>the</strong>temperate zone, or during a period <strong>of</strong> warmth in <strong>the</strong> Antarcticregions, <strong>the</strong>re would have been no such strict limitations to <strong>the</strong>inter-migration <strong>of</strong> <strong>animals</strong>.that floating ice has had some share in <strong>the</strong>It may be held to support <strong>the</strong> viewtransmission <strong>of</strong> fishand amphibia, when -we find that in <strong>the</strong> case <strong>of</strong> <strong>the</strong> narrowtropical sea dividing Borneo from Celebes and <strong>the</strong> Moluccas, noproportionate amount <strong>of</strong> transmission has taken place, butnumerous species, genera, and whole families, terminate abruptlyat what we have o<strong>the</strong>r reasons for believing to be <strong>the</strong> fur<strong>the</strong>stlimits <strong>of</strong> an ancient continent. We can hardly suppose, however,that this mode <strong>of</strong> transmission would have sufficed forsuch groups as tree-frogs, which are inhabitants <strong>of</strong> <strong>the</strong> moretemperate or even warm portions <strong>of</strong> <strong>the</strong> two sou<strong>the</strong>rn lands.Some <strong>of</strong> <strong>the</strong>se cases may perhaps be explained by <strong>the</strong> supposition<strong>of</strong> a considerable extent <strong>of</strong> land in <strong>the</strong> South-Temperate andAntarctic regions now submerged, and by a warm or temperateclimate analogous to that which prevailed in <strong>the</strong> Arctic regionsduring some part <strong>of</strong> <strong>the</strong> Miocene epoch ; while o<strong>the</strong>rs may bedue to cases <strong>of</strong> survival in <strong>the</strong> two areas <strong>of</strong> once wide-spreadgroups, a view supported in <strong>the</strong> case <strong>of</strong> <strong>the</strong> Amphibia by <strong>the</strong>erratic manner in which many <strong>of</strong> <strong>the</strong> groups are spread over<strong>the</strong> globe.From an examination <strong>of</strong> <strong>the</strong> facts presented by <strong>the</strong> variousclasses <strong>of</strong> vertebrates, we are, <strong>the</strong>n, led to <strong>the</strong> conclusion,that <strong>the</strong>re is no evidence <strong>of</strong> a former land-connection be,tween <strong>the</strong> Australian and Neotropical regions ;but that <strong>the</strong>various scattered resemblances in <strong>the</strong>ir natural productions