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The geographical distribution of animals, with a study of the relations ...

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;12 DISTRIBUTION OF ANIMALS. [part i.since <strong>the</strong>y inhabit almost all <strong>the</strong> tropical regions but do notrange more than about 10° beyond <strong>the</strong> sou<strong>the</strong>rn and 12° beyond<strong>the</strong> nor<strong>the</strong>rn tropic, while <strong>the</strong> great bulk <strong>of</strong> <strong>the</strong> species arefound only <strong>with</strong>in an equatorial belt about 30° wide. But as<strong>the</strong>se <strong>animals</strong> are almost exclusively fruit-eaters, <strong>the</strong>ir <strong>distribution</strong>depends as much on vegetation as on temperature ;and thisis. strikingly shown by <strong>the</strong> fact that <strong>the</strong> Semnopi<strong>the</strong>eus schistaceusinhabits <strong>the</strong> Himalayan mountains to a height <strong>of</strong> 11,000feet, where it has been seen leaping among fir-trees loaded <strong>with</strong>snow-wreaths ! Some nor<strong>the</strong>rn <strong>animals</strong> are bounded by <strong>the</strong>iso<strong>the</strong>rmal <strong>of</strong> 32°. Such are <strong>the</strong> polar bear and <strong>the</strong> walrus,which cannot live in a state <strong>of</strong> nature far beyond <strong>the</strong> limits <strong>of</strong><strong>the</strong> frozen ocean ;but as <strong>the</strong>y live in confinement in temperatecountries, <strong>the</strong>ir range is probably limited by o<strong>the</strong>r conditionsthan temperature.We must not <strong>the</strong>refore be too hasty in concluding, that <strong>animals</strong>which we now see confined to a very hot or a very cold climateare incapable <strong>of</strong> living in any o<strong>the</strong>r. <strong>The</strong> tiger was once considereda purely tropical animal, but it inhabits permanently <strong>the</strong>cold plains <strong>of</strong> Manchuria and <strong>the</strong> Amoor, a country <strong>of</strong> an almostarctic winter climate. Few <strong>animals</strong> seem to us more truly inhabitants<strong>of</strong> hot countries than <strong>the</strong> elephants and rhinocerosesyet in Post-tertiary times <strong>the</strong>y roamed over <strong>the</strong> whole <strong>of</strong> <strong>the</strong>nor<strong>the</strong>rn continents to <strong>with</strong>in <strong>the</strong> arctic circle ;and we know that<strong>the</strong> climate was <strong>the</strong>n as cold as it is now, from <strong>the</strong>ir entire bodiesbeing preserved in ice. Some change must recently haveoccurred ei<strong>the</strong>r in <strong>the</strong> climate, soil, or vegetation <strong>of</strong> Nor<strong>the</strong>rnAsia which led to <strong>the</strong> extinction <strong>of</strong> <strong>the</strong>se forerunners <strong>of</strong> existingtropical species ;and we must always bear in mind that similarchanges may have acted upon o<strong>the</strong>r species which we now findrestricted <strong>with</strong>in narrow limits, but which may once have roamedover a wide and varied territory.Valleys and Rivers as Barriers to Mammals.—To <strong>animals</strong> whichthrive best in dry and hilly regions, a broad level and marshyvalley must <strong>of</strong>ten prove an effectual barrier. <strong>The</strong> difference <strong>of</strong>vegetation and <strong>of</strong> insect life,toge<strong>the</strong>r <strong>with</strong> an unhealthy atmosphere,no doubt <strong>of</strong>ten checks migration if it is attempted. Thus

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