The geographical distribution of animals, with a study of the relations ...

The geographical distribution of animals, with a study of the relations ... The geographical distribution of animals, with a study of the relations ...

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—302 ZOOLOGICAL GEOGRAPHY. [part hi.and Malaya to have taken place, we shall perhaps be able toaccount for most of the special affinities they present, with theleast amount of simultaneous elevation of the ocean bed ; whichit must always be remembered, requires a corresponding depressionelsewhere to balance it.Concluding Remarks on the Oriental Region.—We have alreadyso fully discussed the internal and external relations of theseveral sub-regions, that little more need be said. The rich andvaried fauna which inhabited Europe at the dawn of the tertiaryperiod,—as shown by the abundant remains of mammaliawherever suitable deposits of Eocene age have been discovered,proves, that an extensive Palsearctic continent then existed;and the character of the floraisand fauna of the Eocene depositsso completely tropical, that we may be sure there was then nobarrier of climate between it and the Oriental region. At thatearly period the northern plains of Asia were probably underwater, while the great Thibetan plateau and the Himalayan range,had not risen to more than a moderate height, and would havesupported a luxuriant sub*-tropical flora and fauna.The UpperMiocene deposits of northern and central India, and Burmah,agree in their mammalian remains with those of central andsouthern Europe, whileclosely allied forms of elephant, hysena,tapir, rhinoceros, and Chalicotherium have occurred in NorthChina; leading us to conclude that one great fauna thenextended over much of the Oriental and Palsearctic regions.Perim island at the mouth of the Eed Sea, where similarremains are found, probably shows the southern boundary ofthis part of the old Palsearctic region in the Miocene period.Towards the equator there would, of course, be some peculiargroups ; but we can hardly doubt, that, in that wonderful timewhen even the lands that stretched out furthest towards thepole, supported a luxuriant forest vegetation, substantially onefauna ranged over the whole of the great eastern continent of thenorthern hemisphere. During the Pliocene period, however, aprogressive change went on which resulted in the completedifferentiation of the Oriental and Palsearctic faunas. The

chap, xii.] THE ORIENTAL EEGION. 363causes of this change were of two kinds. There was a greatgeographical and physical revolution effected by the elevationof the Himalayas and the Thibetan plateau, and, probably atthe same time, the northward extension of the great Siberianplains. This alone would produce an enormous change ofclimate in all the extra-tropical part of Asia, and inevitablylead to a segregation of the old fauna into tropical and temperate,and a modification of the latter so as to enable it tosupport a climate far more severe than it had previously known.But it is almost certain that, concurrently with this, there wasa change going on of acosmical nature, leading to an alterationof the climate of the northern hemisphere from equable toextreme, and culminating in that period of excessive cold whichdrove the last remnants of the old sub-tropical fauna beyondthe limits of the Palaearctic region. From that time, the Orientaland the Ethiopian regions alone contained the descendants of manyof the most remarkable types which had previously flourishedover all Europe and Asia ; but the early history of these tworegions, and the peculiar equatorial types developed in each,sufficiently separate them, as we have already shown. TheMalayan sub-region is tLat in which characteristic Orientaltypes are now best developed, and where the fundamental contrastof the Oriental, as compared with the Ethiopian andPaleearctic regions, is most distinctly visible.

chap, xii.] THE ORIENTAL EEGION. 363causes <strong>of</strong> this change were <strong>of</strong> two kinds. <strong>The</strong>re was a great<strong>geographical</strong> and physical revolution effected by <strong>the</strong> elevation<strong>of</strong> <strong>the</strong> Himalayas and <strong>the</strong> Thibetan plateau, and, probably at<strong>the</strong> same time, <strong>the</strong> northward extension <strong>of</strong> <strong>the</strong> great Siberianplains. This alone would produce an enormous change <strong>of</strong>climate in all <strong>the</strong> extra-tropical part <strong>of</strong> Asia, and inevitablylead to a segregation <strong>of</strong> <strong>the</strong> old fauna into tropical and temperate,and a modification <strong>of</strong> <strong>the</strong> latter so as to enable it tosupport a climate far more severe than it had previously known.But it is almost certain that, concurrently <strong>with</strong> this, <strong>the</strong>re wasa change going on <strong>of</strong> acosmical nature, leading to an alteration<strong>of</strong> <strong>the</strong> climate <strong>of</strong> <strong>the</strong> nor<strong>the</strong>rn hemisphere from equable toextreme, and culminating in that period <strong>of</strong> excessive cold whichdrove <strong>the</strong> last remnants <strong>of</strong> <strong>the</strong> old sub-tropical fauna beyond<strong>the</strong> limits <strong>of</strong> <strong>the</strong> Palaearctic region. From that time, <strong>the</strong> Orientaland <strong>the</strong> Ethiopian regions alone contained <strong>the</strong> descendants <strong>of</strong> many<strong>of</strong> <strong>the</strong> most remarkable types which had previously flourishedover all Europe and Asia ; but <strong>the</strong> early history <strong>of</strong> <strong>the</strong>se tworegions, and <strong>the</strong> peculiar equatorial types developed in each,sufficiently separate <strong>the</strong>m, as we have already shown. <strong>The</strong>Malayan sub-region is tLat in which characteristic Orientaltypes are now best developed, and where <strong>the</strong> fundamental contrast<strong>of</strong> <strong>the</strong> Oriental, as compared <strong>with</strong> <strong>the</strong> Ethiopian andPaleearctic regions, is most distinctly visible.

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