The geographical distribution of animals, with a study of the relations ...

The geographical distribution of animals, with a study of the relations ... The geographical distribution of animals, with a study of the relations ...

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358 ZOOLOGICAL GEOGRAPHY. [part in.active or extinct, is known in its entire area ; while extensivebeds of coal of tertiary age, in every part of it, prove that it hasbeen subject to repeated submersions, at no distant date geologically.An indication, if not a proof, of still more recent submersionis to be found in the great alluvial valleys which onthe south and south-west extend fully 200 miles inland, whilethey are to a less degree a characteristic feature all round theisland. These swampy plains have been formed by the combinedaction of rivers and tides ; and they point clearly to an immediatelypreceding state of things, when that which is even nowbarely raised above the ocean, was more or less sunk below it.These various indications enable us toclaim, as an admissibleand even probable supposition, that at some epoch during thePliocene period! of geology, Borneo, as we now know it, did notexist ;but was represented by a mountainous island at its presentnorthern extremity, with perhaps a few smaller islets to thesouth. We thus have a clear opening from Java to the SiamesePeninsula ;and as the whole of that sea is less than 100 fathomsdeepj there is no difficulty in supposing an elevation of landconnecting the two together, quite independent of Borneo on theone hand and Sumatra on the other.This union did not probablylast long ; but it was sufficient to allow of the introductioninto Java of the Rhinoceros javanicus, and that group of Indo-Chinese and Himalayan species of mammalia and birds whichit alone possesses. When this ridge had disappeared by subsidence,the next elevation occurred a little more to the east,and produced the union of many islets which, aided by subaerialdenudation, formed the present island of Borneo. It isprobable that this elevation was sufficiently extensive to uniteBorneo for a time with the Malay Peninsula and Sumatra, thushelping to produce that close resemblance of genera and even ofspecies, which these countries exhibit, and obliterating muchot their former speciality, of which, however, we have stillsome traces in the long-nosed monkey and Ptilocerus ofBorneo, and the considerable number of genera both of mammaliaand birds confined to two only out of thethree divisionsof typical Malaya. The subsidence which again divided these

;chap, xii.] THE ORIENTAL REGION. 359countries by arms of the sea rather wider than at present,might have left Banca isolated, as already referred to, with itsproportion of the common fauna to be, in a few instancessubsequently modified.Thus we are enabled to understand how the special relationsof the species of these islands to each other may have beenbrought about To account for their more deep-seated andgeneral zoological features, we must go farther back.Probable Origin of the Malayan Fauna.—The typical Malayanfauna is essentially an equatorial one, and must have beenelaborated in an extensive equatorial area. This ancient landalmost certainly extended northward over the shallow sea as farasthe island of Palawan, the Paracels shoals and even Hainan.To the east, it may at one time have included the Philippinesand Celebes, but not the Moluccas. To the south it was limitedby the deep sea beyond Java. It included all Sumatra and theNicobar islands, and there is every reason to believe that itstretched out also to the west so as to include the central peakof Ceylon, the Maldive isles, and the Cocos islands west ofSumatra. We should then have an area as extensive as SouthAmerica to 15° south latitude, and well calculated to developthat luxuriant fauna and flora which has since spread to theHimalayas. The submergence of the western half of this area(leaving only a fragment in Ceylon) would greatly diminish thenumber of animals and perhaps extinguish some peculiar typesbut the remaining portion would still form a compact and extensivedistrict, twice as large as the peninsula of India, over thewhole of which a uniform Malayan fauna would prevail.Thefirst important change would be the separation of Celebes ;and':his was probably effected by a great subsidence, forming the deepstrait that now divides that island from Borneo. During theprocess Celebesitself was no doubt greatly submerged, leavingonly a few islands in which were preserved that remnant of theancient Malayan fauna that now constitutes one of its moststriking and anomalous features. The Philippine area wouldnext be separated, and perhaps be almost wholly submerged ;or

;chap, xii.] THE ORIENTAL REGION. 359countries by arms <strong>of</strong> <strong>the</strong> sea ra<strong>the</strong>r wider than at present,might have left Banca isolated, as already referred to, <strong>with</strong> itsproportion <strong>of</strong> <strong>the</strong> common fauna to be, in a few instancessubsequently modified.Thus we are enabled to understand how <strong>the</strong> special <strong>relations</strong><strong>of</strong> <strong>the</strong> species <strong>of</strong> <strong>the</strong>se islands to each o<strong>the</strong>r may have beenbrought about To account for <strong>the</strong>ir more deep-seated andgeneral zoological features, we must go far<strong>the</strong>r back.Probable Origin <strong>of</strong> <strong>the</strong> Malayan Fauna.—<strong>The</strong> typical Malayanfauna is essentially an equatorial one, and must have beenelaborated in an extensive equatorial area. This ancient landalmost certainly extended northward over <strong>the</strong> shallow sea as faras<strong>the</strong> island <strong>of</strong> Palawan, <strong>the</strong> Paracels shoals and even Hainan.To <strong>the</strong> east, it may at one time have included <strong>the</strong> Philippinesand Celebes, but not <strong>the</strong> Moluccas. To <strong>the</strong> south it was limitedby <strong>the</strong> deep sea beyond Java. It included all Sumatra and <strong>the</strong>Nicobar islands, and <strong>the</strong>re is every reason to believe that itstretched out also to <strong>the</strong> west so as to include <strong>the</strong> central peak<strong>of</strong> Ceylon, <strong>the</strong> Maldive isles, and <strong>the</strong> Cocos islands west <strong>of</strong>Sumatra. We should <strong>the</strong>n have an area as extensive as SouthAmerica to 15° south latitude, and well calculated to developthat luxuriant fauna and flora which has since spread to <strong>the</strong>Himalayas. <strong>The</strong> submergence <strong>of</strong> <strong>the</strong> western half <strong>of</strong> this area(leaving only a fragment in Ceylon) would greatly diminish <strong>the</strong>number <strong>of</strong> <strong>animals</strong> and perhaps extinguish some peculiar typesbut <strong>the</strong> remaining portion would still form a compact and extensivedistrict, twice as large as <strong>the</strong> peninsula <strong>of</strong> India, over <strong>the</strong>whole <strong>of</strong> which a uniform Malayan fauna would prevail.<strong>The</strong>first important change would be <strong>the</strong> separation <strong>of</strong> Celebes ;and':his was probably effected by a great subsidence, forming <strong>the</strong> deepstrait that now divides that island from Borneo. During <strong>the</strong>process Celebesitself was no doubt greatly submerged, leavingonly a few islands in which were preserved that remnant <strong>of</strong> <strong>the</strong>ancient Malayan fauna that now constitutes one <strong>of</strong> its moststriking and anomalous features. <strong>The</strong> Philippine area wouldnext be separated, and perhaps be almost wholly submerged ;or

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