The geographical distribution of animals, with a study of the relations ...

The geographical distribution of animals, with a study of the relations ... The geographical distribution of animals, with a study of the relations ...

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346 ZOOLOGICAL GEOGRAPHY. [part hi.The foregoing list, although small, contains an assemblage ofspecies which are wholly Oriental in character, and several ofwhich {Tardus, Galeopithecus, Tupaia) are characteristic andhighly peculiar Malayan forms. At the same time these islandsare completely separated from the rest of Malaya by the totalabsence of Semnopithecus, Hylobates, Fclis,Hclarctos, Rhinoceros,Manis, and other groups constantly found in the great Indo-Malay islands and peninsula of Malacca. We find apparentlytwo sets of animals : a more ancient series, represented by thedeer, Galeopithecus, and squirrel, in which the species are distinctfrom any others ; and a more recent series, represented byMacacus cynomolgus, and Viverra tangalunga, identical withcommon Malayan animals. The former indicate the earliestperiod when these volcanic islands were connected with somepart of the Malayan sub-region, and they show that this wasnot geologically remote, since no peculiar generic types havebeen preserved or differentiated. The latter may indicate eitherthe termination of the period of union, or merely the effects ofintroduction by man. The reason why a larger number ofmammalian forms were not introduced and established, wasprobably because the union was effectedonly with some smallislands, and from these communicated to other parts of thearchipelago ; or it may well be that later subsidences extinguishedsome of the forms that had established themselves.Birds of the Philippine Islands.— These have been carefullyinvestigated by Viscount Walden, in a paper read before theZoological Society of London in 1873, and we are thus furnishedwith ample information on the relations of this importantportion of the fauna.The total number of birds known to inhabit the Philippinesis 219, of which 106 are peculiar. If, however, following ourusual plan, we take only the land-birds, we find the numbersto be 159 species, of which 100 are peculiar ;an unusually largeproportion for a group of islands so comparatively near tovarious parts of the Oriental and Australian regions. Thefamilies of birds which are more especially characteristic of theIndo-Malay sub-region are about 28 in number, and examples

—chap, xii.] THE ORIENTAL REGION. 347of all these are found in the Philippines except four, viz., Cinclidse,Phyllornithidse, Eurylsemidae, and Podargidse. The onlyMalayan are, Cacatuidse and Megapodiidae. Yet although thebirds are unmistakably Malayan, as a whole, there are, as inthe mammalia (though in a less degree), marked deficiencies ofmost characteristic Malayan forms. Lord Walden gives a listof no less than 69 genera thus absent ; but it will be sufficienthere to mention such wide-spread and specially Indo-Malaygroupsas,Malacopteron,Eurylcemus, Nydiornis, Arachnotliera, Geocichla,Timalia, Pomatorhinus, Phyllornis, lora, CrinigerEnicurus, Chaptia, Tchitrea, Dendrocitta, Eulahes, Palceornis,Miglyptes, Tiga, and Euplocamus. These deficiencies plainlyshow the isolated character of thethat it has never formed a part of thatPhilippine group, and implyIndo-Malayan extensionof the continent which almost certainly existed when theyPhilippine families which are, otherwise, exclusively Austro-peculiarMalayan fauna was developed ; or that, if it has been sounited, it has been subsequently submerged and broken up tosuch an extent, as to cause the extinction of many of the absenttypes.It appears from Lord "Walden's careful analysis, that 31 of thePhilippine species occur in the Papuan sub-region, and 47 inCelebes ;69 occur also in India, and 75 in Java. This last factiscurious, since Java is the most remote of the Malayan islands,but it is found to arise almost wholly from the birds of thatisland being better known, since only one species, Xantholcemarosea, is confined to the Philippine Islands and Java.The wading and swimming birds are mostly of wide-spreadforms, only 6 out of the 60 species being peculiar to the Philippinearchipelago. Confining ourselves to the land-birds, and combiningseveral of the minutely subdivided genera of Lord Walden'spaper so as to agree with the arrangement adopted in thiswork, we find that there are 112 genera of land-birds representedin the islands. Of these, 50 are eithsr cosmopolitan, ofwide range, or common to the Oriental and Australian regions,and may be put aside as affording few indications of geographicalaffinity. Of the remaining 62 no less than 40 are exclusively

346 ZOOLOGICAL GEOGRAPHY. [part hi.<strong>The</strong> foregoing list, although small, contains an assemblage <strong>of</strong>species which are wholly Oriental in character, and several <strong>of</strong>which {Tardus, Galeopi<strong>the</strong>cus, Tupaia) are characteristic andhighly peculiar Malayan forms. At <strong>the</strong> same time <strong>the</strong>se islandsare completely separated from <strong>the</strong> rest <strong>of</strong> Malaya by <strong>the</strong> totalabsence <strong>of</strong> Semnopi<strong>the</strong>cus, Hylobates, Fclis,Hclarctos, Rhinoceros,Manis, and o<strong>the</strong>r groups constantly found in <strong>the</strong> great Indo-Malay islands and peninsula <strong>of</strong> Malacca. We find apparentlytwo sets <strong>of</strong> <strong>animals</strong> : a more ancient series, represented by <strong>the</strong>deer, Galeopi<strong>the</strong>cus, and squirrel, in which <strong>the</strong> species are distinctfrom any o<strong>the</strong>rs ; and a more recent series, represented byMacacus cynomolgus, and Viverra tangalunga, identical <strong>with</strong>common Malayan <strong>animals</strong>. <strong>The</strong> former indicate <strong>the</strong> earliestperiod when <strong>the</strong>se volcanic islands were connected <strong>with</strong> somepart <strong>of</strong> <strong>the</strong> Malayan sub-region, and <strong>the</strong>y show that this wasnot geologically remote, since no peculiar generic types havebeen preserved or differentiated. <strong>The</strong> latter may indicate ei<strong>the</strong>r<strong>the</strong> termination <strong>of</strong> <strong>the</strong> period <strong>of</strong> union, or merely <strong>the</strong> effects <strong>of</strong>introduction by man. <strong>The</strong> reason why a larger number <strong>of</strong>mammalian forms were not introduced and established, wasprobably because <strong>the</strong> union was effectedonly <strong>with</strong> some smallislands, and from <strong>the</strong>se communicated to o<strong>the</strong>r parts <strong>of</strong> <strong>the</strong>archipelago ; or it may well be that later subsidences extinguishedsome <strong>of</strong> <strong>the</strong> forms that had established <strong>the</strong>mselves.Birds <strong>of</strong> <strong>the</strong> Philippine Islands.— <strong>The</strong>se have been carefullyinvestigated by Viscount Walden, in a paper read before <strong>the</strong>Zoological Society <strong>of</strong> London in 1873, and we are thus furnished<strong>with</strong> ample information on <strong>the</strong> <strong>relations</strong> <strong>of</strong> this importantportion <strong>of</strong> <strong>the</strong> fauna.<strong>The</strong> total number <strong>of</strong> birds known to inhabit <strong>the</strong> Philippinesis 219, <strong>of</strong> which 106 are peculiar. If, however, following ourusual plan, we take only <strong>the</strong> land-birds, we find <strong>the</strong> numbersto be 159 species, <strong>of</strong> which 100 are peculiar ;an unusually largeproportion for a group <strong>of</strong> islands so comparatively near tovarious parts <strong>of</strong> <strong>the</strong> Oriental and Australian regions. <strong>The</strong>families <strong>of</strong> birds which are more especially characteristic <strong>of</strong> <strong>the</strong>Indo-Malay sub-region are about 28 in number, and examples

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