The geographical distribution of animals, with a study of the relations ...

The geographical distribution of animals, with a study of the relations ... The geographical distribution of animals, with a study of the relations ...

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8 DISTRIBUTION OF ANIMALS. [part i.will separate countries the productions of which most widelyand radically differ; while the most recently-depressed seas,or the last-formed mountain ranges, will separate countriesthe productions of which are almost or quite identical. Itwill be evident, therefore, that the study of the distributionof animals and plants may add greatly to our knowledgeof the past history of our globe. It may reveal to us, in amanner which no other evidence can, which are the oldestand most permanent features of the earth's surface, and whichthe newest. It may indicate the existence of islands or continentsnow sunk beneath the ocean, and which have left norecord of their existence save the animal and vegetable productionswhich have migrated to adjacent lands. It thusbecomes an important adjunct to geology, which can rarely domore than determine what lands have been raised above thewaters, under what conditions and at what period; but canseldom ascertain anything of the position or extent of thosewhich have sunk beneath it. Our present study may oftenenable us, not only to say where lands must have recentlydisappeared, but also to form some judgment as to their extent,and the time that has elapsed since their submersion.Having thus briefly sketched the nature and objects of thesubject we have to study, it will be necessary—before enteringon a detailed examination of the zoological features of thedifferent parts of the earth, and of the distribution of the orders,families, and genera of animals— to examine certain preliminaryfacts and principles essential for our guidance. We must firstinquire what are the powers of multiplication and dispersal ofthe various groups of animals, and the nature of the barriersthat most effectually limit their range. We have then toconsider the effects of changes in physical geography and inclimate ;to examine the nature and extent of such changes ashave been known to occur ; to determine what others are possibleor probable ; and to ascertain the various modes in which suchchanges affect the structure, the distribution, or the very existenceof animals.

3chap, i.] INTRODUCTORY. 9Two subjects of a different nature must next engage ourattention. We have to deal with two vast masses of facts,each involving countlessdetails, and requiring subdivision andgrouping to be capable of intelligible treatment. All the continentsand their chief subdivisions, and all the more importantislands of the globe, have to be compared as regards their variousanimal forms. To do this effectively we require a naturaldivision of the earth especially adapted to our purpose ; and weshall have to discuss at some length the reasons for the particularsystem adopted,—a discussion which must to some extentanticipate and summarize the conclusions of the whole work.We have also to deal with many hundreds of families and manythousands of genera ofanimals, and here too a true and naturalclassification is of great importance. We must therefore give aconnected view of the classification adopted in the variousclasses of animals dealt with.And lastly, as the existing distribution of animals is theresult and outcome ofall preceding changes of the earth and ofitsinhabitants, we require as much knowledge as we can get ofthe animals of each country during past geological epochs, inorder to interpret the facts we shall accumulate. We shall,therefore, enter upon a somewhat detailed sketch of the variousforms of extinct animals that have lived upon the earthduringthe Tertiary period ; discuss their migrations at various epochs,the changes of physical geography that they imply, and theextent to which they enable us to determine the birthplaceof certain families and genera.The preliminary studies above enumerated will, it is believed,enable us to see the bearing of many facts in the distribution ofanimals that would otherwise be insoluble problems ;and, whatis hardly less valuable, will teach us to estimate the comparativeimportance of the various groups of animals, and to avoidthe common error of cutting the gordian knot of each difficultyby vast hypothetical changes in existing continents and oceans—probably the most permanent features of our globe.Vol. I—

8 DISTRIBUTION OF ANIMALS. [part i.will separate countries <strong>the</strong> productions <strong>of</strong> which most widelyand radically differ; while <strong>the</strong> most recently-depressed seas,or <strong>the</strong> last-formed mountain ranges, will separate countries<strong>the</strong> productions <strong>of</strong> which are almost or quite identical. Itwill be evident, <strong>the</strong>refore, that <strong>the</strong> <strong>study</strong> <strong>of</strong> <strong>the</strong> <strong>distribution</strong><strong>of</strong> <strong>animals</strong> and plants may add greatly to our knowledge<strong>of</strong> <strong>the</strong> past history <strong>of</strong> our globe. It may reveal to us, in amanner which no o<strong>the</strong>r evidence can, which are <strong>the</strong> oldestand most permanent features <strong>of</strong> <strong>the</strong> earth's surface, and which<strong>the</strong> newest. It may indicate <strong>the</strong> existence <strong>of</strong> islands or continentsnow sunk beneath <strong>the</strong> ocean, and which have left norecord <strong>of</strong> <strong>the</strong>ir existence save <strong>the</strong> animal and vegetable productionswhich have migrated to adjacent lands. It thusbecomes an important adjunct to geology, which can rarely domore than determine what lands have been raised above <strong>the</strong>waters, under what conditions and at what period; but canseldom ascertain anything <strong>of</strong> <strong>the</strong> position or extent <strong>of</strong> thosewhich have sunk beneath it. Our present <strong>study</strong> may <strong>of</strong>tenenable us, not only to say where lands must have recentlydisappeared, but also to form some judgment as to <strong>the</strong>ir extent,and <strong>the</strong> time that has elapsed since <strong>the</strong>ir submersion.Having thus briefly sketched <strong>the</strong> nature and objects <strong>of</strong> <strong>the</strong>subject we have to <strong>study</strong>, it will be necessary—before enteringon a detailed examination <strong>of</strong> <strong>the</strong> zoological features <strong>of</strong> <strong>the</strong>different parts <strong>of</strong> <strong>the</strong> earth, and <strong>of</strong> <strong>the</strong> <strong>distribution</strong> <strong>of</strong> <strong>the</strong> orders,families, and genera <strong>of</strong> <strong>animals</strong>— to examine certain preliminaryfacts and principles essential for our guidance. We must firstinquire what are <strong>the</strong> powers <strong>of</strong> multiplication and dispersal <strong>of</strong><strong>the</strong> various groups <strong>of</strong> <strong>animals</strong>, and <strong>the</strong> nature <strong>of</strong> <strong>the</strong> barriersthat most effectually limit <strong>the</strong>ir range. We have <strong>the</strong>n toconsider <strong>the</strong> effects <strong>of</strong> changes in physical geography and inclimate ;to examine <strong>the</strong> nature and extent <strong>of</strong> such changes ashave been known to occur ; to determine what o<strong>the</strong>rs are possibleor probable ; and to ascertain <strong>the</strong> various modes in which suchchanges affect <strong>the</strong> structure, <strong>the</strong> <strong>distribution</strong>, or <strong>the</strong> very existence<strong>of</strong> <strong>animals</strong>.

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