The geographical distribution of animals, with a study of the relations ...
The geographical distribution of animals, with a study of the relations ... The geographical distribution of animals, with a study of the relations ...
4 DISTRIBUTION OF ANIMALS. [part i.the soil, the aspect, or the vegetation, was adapted to them orthe reverse. The marshes, the heaths, the woods and forests,the chalky downs, the rocky mountains, had each their peculiarinhabitants, which reappeared again and again as we came totracts of country suitable for them. But as we got further awaywe began to find that localities very similar to those we hadleft behind were inhabited by a somewhat different set of species;and this difference increased with distance, notwithstandingthat almost identical external conditions might be often metwith. The first class of changes is that of stations ; the secondthat of habitats. The one is a local, the other a geograiihicalphenomenon. The whole area over which a particular animalis found may consist of any number of stations, but rarely ofmore than one habitat.Stations, however, are often so extensiveas to include the entire range of many species. Such are thegreat seas and oceans, the Siberian or the Amazonian forests,the North African deserts, the Andean or the Himalayanhighlands.There is yet another difference in the nature of the changewe have been considering. The new animals which we meetwith as we travel in any direction from our starting point, aresome of them very much like those we have left behind us,and can be at once referred to familiar types ; while othersare altogether unlike anything we have seen at home. Whenwe reach the Alps we find another kind of squirrel, in SouthernItaly a distinct mole, in Southern Europe fresh warblersand unfamiliar buntings. We meet also with totally newforms ;as the glutton and the snowy owl in Northern, the genetand the hoopoe in Southern, and the saiga antelope andcollared pratincole in Eastern Europe. The first series areexamples of what are termed representative species, the secondof distinct groups or types of animals. The one represents acomparatively recent modification, and an origin in or near thelocality where it occurs ; the other is a result of very ancientchanges both organic and inorganic, and is connected with someof theto discuss.most curious and difficult of the problems we shall have
chap, i.] INTRODUCTORY.Having thus defined our subject, let us glance at the opinionsthat have generally prevailed as to the nature and causes ofthe phenomena presented by the geographical distribution ofanimals.It was long thought, and is still a popular notion, that themanner in which the various kinds of animals are dispersedover the globe is almost wholly due to diversities of climate andof vegetation. There is indeed much to favour this belief. Thearctic regions are strongly characterised by their white bearsand foxes, their reindeer, ermine, and walruses, their whiteptarmigan, owls, and falcons ;the temperate zone has its foxesand wolves, its rabbits, sheep, beavers, and marmots, its sparrowsand its song birds ;while tropical regions alone produce apes andelephants, parrots and peacocks, and a thousand strange quadrupedsand brilliant birds which arefound nowhere in the coolerregions. So the camel, the gazelle and the ostrich live in thedesert ; the bison on the prairie ; the tapir, the deer, and thejaguar in forests. Mountains and marshes, plains and rockyprecipices, have each their animal inhabitants ;•be thought, in the absenceand it might wellof accurate inquiry, that these andother differences would sufficiently explain why most of theregions and countriesinto which the earth is popularly dividedshould have certain animals peculiar to them and should wantothers which are elsewhere abundant.A more detailed and accurate knowledge of the productions ofdifferent portions of the earth soon showed that this explanationwas quite insufficient ; for it was found that countries exceedinglysimilar in climate and all physicalvery distinct animal populations.features may yet haveThe equatorial parts of Africaand South America, for example, are very similar in climateand are both covered with luxuriant forests, yet their animal lifeis widely different; elephants, apes, leopards, guinea-fowlsand touracos in the one, are replaced by tapirs, prehensiletailedmonkeys, jaguars, curassows and toucans in the other.Again, parts of South Africa and Australia are wonderfullysimilar in their soil and climate;yet one has lions, antelopes,zebras and giraffes ; the other only kangaroos, wombats, phalan-
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4 DISTRIBUTION OF ANIMALS. [part i.<strong>the</strong> soil, <strong>the</strong> aspect, or <strong>the</strong> vegetation, was adapted to <strong>the</strong>m or<strong>the</strong> reverse. <strong>The</strong> marshes, <strong>the</strong> heaths, <strong>the</strong> woods and forests,<strong>the</strong> chalky downs, <strong>the</strong> rocky mountains, had each <strong>the</strong>ir peculiarinhabitants, which reappeared again and again as we came totracts <strong>of</strong> country suitable for <strong>the</strong>m. But as we got fur<strong>the</strong>r awaywe began to find that localities very similar to those we hadleft behind were inhabited by a somewhat different set <strong>of</strong> species;and this difference increased <strong>with</strong> distance, not<strong>with</strong>standingthat almost identical external conditions might be <strong>of</strong>ten met<strong>with</strong>. <strong>The</strong> first class <strong>of</strong> changes is that <strong>of</strong> stations ; <strong>the</strong> secondthat <strong>of</strong> habitats. <strong>The</strong> one is a local, <strong>the</strong> o<strong>the</strong>r a geograiihicalphenomenon. <strong>The</strong> whole area over which a particular animalis found may consist <strong>of</strong> any number <strong>of</strong> stations, but rarely <strong>of</strong>more than one habitat.Stations, however, are <strong>of</strong>ten so extensiveas to include <strong>the</strong> entire range <strong>of</strong> many species. Such are <strong>the</strong>great seas and oceans, <strong>the</strong> Siberian or <strong>the</strong> Amazonian forests,<strong>the</strong> North African deserts, <strong>the</strong> Andean or <strong>the</strong> Himalayanhighlands.<strong>The</strong>re is yet ano<strong>the</strong>r difference in <strong>the</strong> nature <strong>of</strong> <strong>the</strong> changewe have been considering. <strong>The</strong> new <strong>animals</strong> which we meet<strong>with</strong> as we travel in any direction from our starting point, aresome <strong>of</strong> <strong>the</strong>m very much like those we have left behind us,and can be at once referred to familiar types ; while o<strong>the</strong>rsare altoge<strong>the</strong>r unlike anything we have seen at home. Whenwe reach <strong>the</strong> Alps we find ano<strong>the</strong>r kind <strong>of</strong> squirrel, in Sou<strong>the</strong>rnItaly a distinct mole, in Sou<strong>the</strong>rn Europe fresh warblersand unfamiliar buntings. We meet also <strong>with</strong> totally newforms ;as <strong>the</strong> glutton and <strong>the</strong> snowy owl in Nor<strong>the</strong>rn, <strong>the</strong> genetand <strong>the</strong> hoopoe in Sou<strong>the</strong>rn, and <strong>the</strong> saiga antelope andcollared pratincole in Eastern Europe. <strong>The</strong> first series areexamples <strong>of</strong> what are termed representative species, <strong>the</strong> second<strong>of</strong> distinct groups or types <strong>of</strong> <strong>animals</strong>. <strong>The</strong> one represents acomparatively recent modification, and an origin in or near <strong>the</strong>locality where it occurs ; <strong>the</strong> o<strong>the</strong>r is a result <strong>of</strong> very ancientchanges both organic and inorganic, and is connected <strong>with</strong> some<strong>of</strong> <strong>the</strong>to discuss.most curious and difficult <strong>of</strong> <strong>the</strong> problems we shall have