The geographical distribution of animals, with a study of the relations ...
The geographical distribution of animals, with a study of the relations ... The geographical distribution of animals, with a study of the relations ...
;230 ZOOLOGICAL GEOGRAPHY. [part hi.the southern latitude and warm climate in which theseinsectswere mostly collected, and the proximity to Formosa and theMalay Islands compared with the enormous distance fromEurope, this shows as much Palaearctic affinity as can beexpected. In the Palaearctic region the group is only plentifulin the southern parts of Europe, which is cutoff by the coldplateau of Thibet from all direct communication with Japan;while in the Oriental region it everywhere abounds and is, infact, one of the most conspicuous and dominant families ofColeoptera.The Longicorns collected by Mr. Lewis have been describedby Mr. Bates in the Annals of Natural History for 1873.number of species now known from Japan isThe107, belonging tosixty-four genera. The most important genera are Lcptura,'Clytanthus, Monohammus, Praonctha, Exoccntrus, Glenea, andOberea. There are twenty-one tropical genera, and sevenpeculiar to Japan, leaving thirty-six either Palaearctic or of verywide range. A number of the genera are Oriental and Malayan,and many characteristic European genera seem to be absentbut it is certaiu that not half the Japanese Longicorns are yetknown, and many of these gaps will doubtless be filled up whenthe more northern islands are explored.The Phytophaga, described by Mr. Baly, appear to have aconsiderable preponderance of tropical Oriental forms.A considerable collection of Hymenoptera formed by Mr.Lewis have been described by Mr. Frederick Smith ;and exhibitthe interesting result, that while the bees and wasps are decidedlyof tropical and Oriental forms, the Tenthredinidae and Ichneumonidaeare as decidedly Pala?arctic, " the generalaspect of thecollection being that of a European one, only a single exoticform being found among them."Remarks on the General Character of the Fauna of Japan —From a general view of the phenomena of distribution we feeljustified in placing Japan in the Palaearctic region ; althoughsome tropical groups, especially of reptiles and insects, havelargely occupied its southern portions ;and these same groupshave in many cases spread into Northern China, beyond the
chap, x.] THE PAL^EARCTIC REGION 231usual dividing line of the Palsearctic and Oriental regions. Thecauses of such a phenomenon are not difficult to conceive. Evennow, that portion of the Palsearctic region between WesternAsia and Japan is, for the most part, a bleak and inhospitableregion, abounding in desert plateaus, and with a rigorous climateeven in its most favoured districts, and can, therefore, supportbut a scanty population of snakes, and of such groups ofinsects as require flowers, forests, or a considerable period ofwarm summer weather ; and it is precisely these which arerepresented in Japan and North China by tropical forms.Wemust also consider, that during the Glacial epoch this wholeregion would have become stillless productive, and that, as thesouthern limit of the ice retired northward, it would be followedup by many tropical forms along with such as had been drivensouth by its advance, and had survived to return to theirnorthern homes.It isalso evident that Japan has a more equable and probablymoister climate than the opposite shores of China, and has alsoa very different geological character, being rocky and broken,often volcanic, and supporting a rich,varied, and peculiar vegetation.It would thus be well adapted to support all the morehardy denizens of the tropics which might at various timesreach it, while it might not be so well adapted for the moreboreal forms from Mongolia or Siberia. The fact that a mixtureof such forms occurs there, is then, little to be wondered at, butwe may rather marvel that they are not more predominant, andthat even in the extreme south, the most abundant forms ofmammal, bird, and insect, are modifications of familiar Palaearctictypes. The fact clearly indicates that the former land connectionsof Japan with the continent have been in a northerlyrather than in a southerly direction, and that the tropical immigrantshave had difficulties to contend with, and have found theland already fairly stocked with northern aborigines inevery class and order of animals.almostGeneral Conclusions as to the Fauna of the Palccarctic Region.—Fromthe account that has now been given of the fauna
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chap, x.] THE PAL^EARCTIC REGION 231usual dividing line <strong>of</strong> <strong>the</strong> Palsearctic and Oriental regions. <strong>The</strong>causes <strong>of</strong> such a phenomenon are not difficult to conceive. Evennow, that portion <strong>of</strong> <strong>the</strong> Palsearctic region between WesternAsia and Japan is, for <strong>the</strong> most part, a bleak and inhospitableregion, abounding in desert plateaus, and <strong>with</strong> a rigorous climateeven in its most favoured districts, and can, <strong>the</strong>refore, supportbut a scanty population <strong>of</strong> snakes, and <strong>of</strong> such groups <strong>of</strong>insects as require flowers, forests, or a considerable period <strong>of</strong>warm summer wea<strong>the</strong>r ; and it is precisely <strong>the</strong>se which arerepresented in Japan and North China by tropical forms.Wemust also consider, that during <strong>the</strong> Glacial epoch this wholeregion would have become stillless productive, and that, as <strong>the</strong>sou<strong>the</strong>rn limit <strong>of</strong> <strong>the</strong> ice retired northward, it would be followedup by many tropical forms along <strong>with</strong> such as had been drivensouth by its advance, and had survived to return to <strong>the</strong>irnor<strong>the</strong>rn homes.It isalso evident that Japan has a more equable and probablymoister climate than <strong>the</strong> opposite shores <strong>of</strong> China, and has alsoa very different geological character, being rocky and broken,<strong>of</strong>ten volcanic, and supporting a rich,varied, and peculiar vegetation.It would thus be well adapted to support all <strong>the</strong> morehardy denizens <strong>of</strong> <strong>the</strong> tropics which might at various timesreach it, while it might not be so well adapted for <strong>the</strong> moreboreal forms from Mongolia or Siberia. <strong>The</strong> fact that a mixture<strong>of</strong> such forms occurs <strong>the</strong>re, is <strong>the</strong>n, little to be wondered at, butwe may ra<strong>the</strong>r marvel that <strong>the</strong>y are not more predominant, andthat even in <strong>the</strong> extreme south, <strong>the</strong> most abundant forms <strong>of</strong>mammal, bird, and insect, are modifications <strong>of</strong> familiar Palaearctictypes. <strong>The</strong> fact clearly indicates that <strong>the</strong> former land connections<strong>of</strong> Japan <strong>with</strong> <strong>the</strong> continent have been in a nor<strong>the</strong>rlyra<strong>the</strong>r than in a sou<strong>the</strong>rly direction, and that <strong>the</strong> tropical immigrantshave had difficulties to contend <strong>with</strong>, and have found <strong>the</strong>land already fairly stocked <strong>with</strong> nor<strong>the</strong>rn aborigines inevery class and order <strong>of</strong> <strong>animals</strong>.almostGeneral Conclusions as to <strong>the</strong> Fauna <strong>of</strong> <strong>the</strong> Palccarctic Region.—From<strong>the</strong> account that has now been given <strong>of</strong> <strong>the</strong> fauna