The geographical distribution of animals, with a study of the relations ...
The geographical distribution of animals, with a study of the relations ... The geographical distribution of animals, with a study of the relations ...
;228 ZOOLOGICAL GEOGRAPHY. [part hi.Coleoptera.—The beetles of Japan decidedly exhibit a mixtureof tropical forms with others truly Palsearctic, and it has beenwith some naturalists a matter of doubt whether the southern andbest known portion of the islands should not be joined to theOriental region. An important addition to our knowledge ofthe insects of this country has recently been made by Mr. GeorgeLewis, and a portion of his collections have been described byvarious entomologists in the Transactions of the EntomologicalSociety of London. As the question is one of considerable interestwe shall give a summary of the results fairly deduciblefrom what is now known of the entomology of Japan ;and itmust be remembered that almost all our collections come fromthe southern districts, in what is almost a sub-tropical climateso that if we find a considerable proportion of Palsearctic forms,we may be pretty sure that the preponderance will be muchgreater a little further north.Of Carabidae Mr. Bates enumerates 244 speciesbelonging to84 genera, and by comparing these with the Coleoptera of atract of about equal extent in western Europe, he concludes thatthere is little similarity, and that the cases of affinity to the formsof eastern tropical Asia preponderate. By comparing his generawith the distributions as given in Gemminger and Harold'sCatalogue, a somewhat different result is arrived at. Leavingout the generic types altogether peculiar to Japan, and also thoseo-enera of such world-wide distribution that they afford no clearindications for our purpose, it appears that no less than twentytwogenera, containing seventy-four of the Japanese species, areeither exclusively Palsearctic, Palsearctic and Nearctic, or highlycharacteristic of the Palsearctic region ; then come thirteen generacontaining eighty-seven of the species which have a very widedistribution, but are also Palsearctic : we next have seventeen^enera containing twenty-four of the Japanese species which aredecidedly Oriental and tropical. Here then the fair comparisonis between the twenty-two genera and seventy-four species whoseaffinities are clearly Palsearctic or at least north temperate, andseventeen genera with twenty-four species which are Asiaticand tropical; and this seems to prove that, although South
chap, x.] THE PAL^ARCTIC REGION. 229Japan (like North China) has a considerable infusion of tropicalforms, there is a preponderating substratum of Palrearctic forms,which clearly indicate the true position of the islandsin zoologicalgeography. There are also a few cases of what may bewhich show that Japan, like manycalled eccentric distribution ;other island-groups, has served as a kind of refuge in whichdying-out forms continue to maintain themselves.These, whichare worthy of notice, are as follows : Orthotrichus (1 sp.) hasthe only other species in Egypt ; Trechichus (1 sp.) has twoother species, of which one inhabits Madeira, the other theSouthern United States ;Perileptus (1 sp.) has two other species,of which one inhabits Bourbon, the other West Europe ; andlastly, Crepidogaster (1 sp.) has the other known species inSouth Africa.These cases diminish the value of the indicationsafforded by some of the Japanese forms, whose only allies aresingle species in various remote parts of the Oriental region.The Staphylinidse have been describedby Dr. Sharp, and hislist exhibits a great preponderance of north temperate, or cosmopolitanforms, with a few which are decidedly tropical. ThePselaphidae and Scydmenidre, also described by Dr. Sharp,exhibit, according to that gentleman, " even a greater resemblanceto those of North America than to those of Europe," but he saysnothing of any tropical affinities. The water-beetles are alleither Paleearctic or of wide distribution.The Lucanidse{Gemm. and Har. Cat, 1868) exhibit an interminglingof Palaearctic and Oriental genera.The Cetoniidse {Gemm. and Har. Gat. 1869) show, for NorthChina and Japan, three Oriental to two Palaearctic genera.The Buprestidse collected by Mr. Lewis have been describedby Mr. Edward Saunders in the Journal of the Linnman Society,vol. xi. p. 509. The collection consisted of thirty-six speciesbelonging to fourteen genera. No less than thirteen of theseare known also from India and the Malay Islands ; nine fromEurope ; seven from Africa ; six from America, and four fromChina.In six of the genera the Japanese species are said to beallied to those of the Oriental region ;while in three they areallied to European forms, and in two to American.Considering
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chap, x.] THE PAL^ARCTIC REGION. 229Japan (like North China) has a considerable infusion <strong>of</strong> tropicalforms, <strong>the</strong>re is a preponderating substratum <strong>of</strong> Palrearctic forms,which clearly indicate <strong>the</strong> true position <strong>of</strong> <strong>the</strong> islandsin zoologicalgeography. <strong>The</strong>re are also a few cases <strong>of</strong> what may bewhich show that Japan, like manycalled eccentric <strong>distribution</strong> ;o<strong>the</strong>r island-groups, has served as a kind <strong>of</strong> refuge in whichdying-out forms continue to maintain <strong>the</strong>mselves.<strong>The</strong>se, whichare worthy <strong>of</strong> notice, are as follows : Orthotrichus (1 sp.) has<strong>the</strong> only o<strong>the</strong>r species in Egypt ; Trechichus (1 sp.) has twoo<strong>the</strong>r species, <strong>of</strong> which one inhabits Madeira, <strong>the</strong> o<strong>the</strong>r <strong>the</strong>Sou<strong>the</strong>rn United States ;Perileptus (1 sp.) has two o<strong>the</strong>r species,<strong>of</strong> which one inhabits Bourbon, <strong>the</strong> o<strong>the</strong>r West Europe ; andlastly, Crepidogaster (1 sp.) has <strong>the</strong> o<strong>the</strong>r known species inSouth Africa.<strong>The</strong>se cases diminish <strong>the</strong> value <strong>of</strong> <strong>the</strong> indicationsafforded by some <strong>of</strong> <strong>the</strong> Japanese forms, whose only allies aresingle species in various remote parts <strong>of</strong> <strong>the</strong> Oriental region.<strong>The</strong> Staphylinidse have been describedby Dr. Sharp, and hislist exhibits a great preponderance <strong>of</strong> north temperate, or cosmopolitanforms, <strong>with</strong> a few which are decidedly tropical. <strong>The</strong>Pselaphidae and Scydmenidre, also described by Dr. Sharp,exhibit, according to that gentleman, " even a greater resemblanceto those <strong>of</strong> North America than to those <strong>of</strong> Europe," but he saysnothing <strong>of</strong> any tropical affinities. <strong>The</strong> water-beetles are allei<strong>the</strong>r Paleearctic or <strong>of</strong> wide <strong>distribution</strong>.<strong>The</strong> Lucanidse{Gemm. and Har. Cat, 1868) exhibit an intermingling<strong>of</strong> Palaearctic and Oriental genera.<strong>The</strong> Cetoniidse {Gemm. and Har. Gat. 1869) show, for NorthChina and Japan, three Oriental to two Palaearctic genera.<strong>The</strong> Buprestidse collected by Mr. Lewis have been describedby Mr. Edward Saunders in <strong>the</strong> Journal <strong>of</strong> <strong>the</strong> Linnman Society,vol. xi. p. 509. <strong>The</strong> collection consisted <strong>of</strong> thirty-six speciesbelonging to fourteen genera. No less than thirteen <strong>of</strong> <strong>the</strong>seare known also from India and <strong>the</strong> Malay Islands ; nine fromEurope ; seven from Africa ; six from America, and four fromChina.In six <strong>of</strong> <strong>the</strong> genera <strong>the</strong> Japanese species are said to beallied to those <strong>of</strong> <strong>the</strong> Oriental region ;while in three <strong>the</strong>y areallied to European forms, and in two to American.Considering