The geographical distribution of animals, with a study of the relations ...
The geographical distribution of animals, with a study of the relations ... The geographical distribution of animals, with a study of the relations ...
168 DISTRIBUTION OF EXTINCT ANIMALS. [part II.and recognisable.It is only when we go further back still, intothe Palseozoic formations, that the insect forms begin to show thatgeneralization of type which renders it impossible to classifythem in any existing groups. Yet even in the coal formation ofNova Scotia and Durham, the fossil insects are said by competententomologists to be " allied to Ephemera" " near Blatta,"Phasmidce ; "" nearand in deposits of the same age at Saarbruck nearTreves, a well-preserved wing of a grasshopper or locust has beenfound, as well as a beetle referred to the Scarabeidse. Moreremarkable, however, is the recent discovery in the carboniferousshales of Belgium, of the clearly-defined wing of a large moth(Breyeria borinensis), closely resembling some of the Saturniidae ;so that we have now all the chief orders of Insects—includingthose supposed to be the most highly developed and the mostrecent—well represented at this very remote epoch. Even theoldest insects, from the Devonian rocks of North America, canmostly be classed as Neuroptera or Myriapoda, but appear toform new families.We may consider it, therefore, as proved, that many of thelarger and more important genera of insects date back to thebeginning of the Tertiary period, or perhaps beyond it ;but thefamily types are far older, and must have been differentiated veryearly in the Secondary period, while some of them perhaps goback to Palseozoic times. The great comparative antiquity ofthe genera is however the important fact for us, and we shallhave occasion often to refer to it, in endeavouring to ascertainthe true bearing of the facts of insect distribution, as elucidatingor invalidating the conclusions arrived at from a study of thedistribution of the higher animals.Antiquity of the Genera of Land and Fresh-WaterShells.The remains of land and fresh-water shells are not much morefrequent than those of insects. Like them, too, their forms arevery stable, continuing unchanged through several geological
—chap, viii.] ANTIQUITY OF LAND SHELLS. 169periods. In the Pliocene and Miocene formations, most of theshells are very similar to living species, and some are quite identical.In the Eocene we meet with ordinary forms of the generaHelix, Clausilia, Pupa, Bulimus, Glandina, Cyclostoma, Megalostoma,Planorbis, Paludina and Limncea, some resembling Europeanspecies, others more like tropical forms. A British Eocenespecies of Helix is still living in Texas ;and in the South of Franceare found species of the Brazilian sub-genera Megaspira andAnastoma. In the secondary formation no true land shells havebeen found, but fresh water shells are tolerably abundant, andalmost all are still of living forms. In the Wealden (LowerCretaceous) and Purbeck (Upper Oolite) are found Unio, Melania,Paludina, Planorbis, and Limncea ; while the last named genusoccurs even in the Lias.The notion that land shells were really not in existence duringthe secondary period is, however, proved to be erroneous by thestartling discovery, in the Palaeozoic coal measures of Nova Scotia,of two species of Helicidae, both of living genera Pupa vetusta,and Zonites priscus. They have been found in the hollow trunkof a Sigillaria, and in great quantities in a bed full of Stigmarianrootlets. The most minute examination detects no importantdifferences of form or of microscopic structure, between theseshells and living species of the same genera !These mollusca werethe contemporaries of Labyrinthodonts and strange Ganoid fishes,which formed almost the whole vertebrate fauna. This unexpecteddiscovery renders it almost certain, that numbers of otherexisting genera, of which we have found no traces, lived withthese two through the whole secondary period ; and we are thusobliged to assume as a probability, that any particular genus haslived through a long succession of geological ages. In estimatingthe importance of any peculiarities or anomalies in thegeographical distribution of land shells as compared with thehigher vertebrates, we shall, therefore, have to keep this possible,and even probable high antiquity, constantly in mind.We have now concluded our sketch of Tertiary Palaeontologyas a preparation for the intelligent study of the GeographicalVol. I.—13
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168 DISTRIBUTION OF EXTINCT ANIMALS. [part II.and recognisable.It is only when we go fur<strong>the</strong>r back still, into<strong>the</strong> Palseozoic formations, that <strong>the</strong> insect forms begin to show thatgeneralization <strong>of</strong> type which renders it impossible to classify<strong>the</strong>m in any existing groups. Yet even in <strong>the</strong> coal formation <strong>of</strong>Nova Scotia and Durham, <strong>the</strong> fossil insects are said by competententomologists to be " allied to Ephemera" " near Blatta,"Phasmidce ; "" nearand in deposits <strong>of</strong> <strong>the</strong> same age at Saarbruck nearTreves, a well-preserved wing <strong>of</strong> a grasshopper or locust has beenfound, as well as a beetle referred to <strong>the</strong> Scarabeidse. Moreremarkable, however, is <strong>the</strong> recent discovery in <strong>the</strong> carboniferousshales <strong>of</strong> Belgium, <strong>of</strong> <strong>the</strong> clearly-defined wing <strong>of</strong> a large moth(Breyeria borinensis), closely resembling some <strong>of</strong> <strong>the</strong> Saturniidae ;so that we have now all <strong>the</strong> chief orders <strong>of</strong> Insects—includingthose supposed to be <strong>the</strong> most highly developed and <strong>the</strong> mostrecent—well represented at this very remote epoch. Even <strong>the</strong>oldest insects, from <strong>the</strong> Devonian rocks <strong>of</strong> North America, canmostly be classed as Neuroptera or Myriapoda, but appear t<strong>of</strong>orm new families.We may consider it, <strong>the</strong>refore, as proved, that many <strong>of</strong> <strong>the</strong>larger and more important genera <strong>of</strong> insects date back to <strong>the</strong>beginning <strong>of</strong> <strong>the</strong> Tertiary period, or perhaps beyond it ;but <strong>the</strong>family types are far older, and must have been differentiated veryearly in <strong>the</strong> Secondary period, while some <strong>of</strong> <strong>the</strong>m perhaps goback to Palseozoic times. <strong>The</strong> great comparative antiquity <strong>of</strong><strong>the</strong> genera is however <strong>the</strong> important fact for us, and we shallhave occasion <strong>of</strong>ten to refer to it, in endeavouring to ascertain<strong>the</strong> true bearing <strong>of</strong> <strong>the</strong> facts <strong>of</strong> insect <strong>distribution</strong>, as elucidatingor invalidating <strong>the</strong> conclusions arrived at from a <strong>study</strong> <strong>of</strong> <strong>the</strong><strong>distribution</strong> <strong>of</strong> <strong>the</strong> higher <strong>animals</strong>.Antiquity <strong>of</strong> <strong>the</strong> Genera <strong>of</strong> Land and Fresh-WaterShells.<strong>The</strong> remains <strong>of</strong> land and fresh-water shells are not much morefrequent than those <strong>of</strong> insects. Like <strong>the</strong>m, too, <strong>the</strong>ir forms arevery stable, continuing unchanged through several geological