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The geographical distribution of animals, with a study of the relations ...

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chap, viii.] MESOZOIC MAMMALIA. 159young born each year in <strong>the</strong> world are probably as some millionsto one ; and it is very easily conceivable that in a thousandyears <strong>the</strong> former might, under pressure <strong>of</strong> rapidly changing conditions,become modified into a distinct species, while <strong>the</strong> latter,not <strong>of</strong>fering enough favourable variations to effect a suitableadaptation, would become extinct. We must also remember<strong>the</strong> extreme specialization <strong>of</strong> many <strong>of</strong> <strong>the</strong> large <strong>animals</strong> thathave become extinct—a specializationwhich would necessarilyrender modification in any new direction difficult, since <strong>the</strong> inheritedtendency <strong>of</strong> variation would probably be to increase <strong>the</strong>specialization in <strong>the</strong> same directionswhich had heret<strong>of</strong>ore beenbeneficial. If to <strong>the</strong>se two causes we add <strong>the</strong> difficulty <strong>of</strong>obtaining sufficient food for such large <strong>animals</strong>, and perhaps <strong>the</strong>injurious effects <strong>of</strong> changes <strong>of</strong> climate, we shall not find it difficultto understand how such a vast physical revolution as <strong>the</strong>Glacial epoch, <strong>with</strong> its attendant phenomena <strong>of</strong> elevations andsubsidences, icy winds, and sudden floods by <strong>the</strong> bursting <strong>of</strong> lakebarriers, might have led to <strong>the</strong> total extinction <strong>of</strong> a vast number<strong>of</strong> <strong>the</strong> most bulky forms <strong>of</strong> mammalia, while <strong>the</strong> less bulkywere able to survive, ei<strong>the</strong>rby greater hardiness <strong>of</strong> constitutionor bv becoming more or less modified. <strong>The</strong> result is apparent intrie comparatively small or moderate size <strong>of</strong> <strong>the</strong> species constituting<strong>the</strong> temperate fauna, in all parts <strong>of</strong> <strong>the</strong> globe.It is much to be regretted that no mammalian remains <strong>of</strong> earlierdate have been found in Australia, as we should <strong>the</strong>n see ifit isreally <strong>the</strong> case that marsupials have always formed its highesttype <strong>of</strong> mammalian life. At present its fossil fauna is chieflyinteresting to <strong>the</strong> zoologist, but throws little light on <strong>the</strong> past<strong>relations</strong> <strong>of</strong> this isolated country <strong>with</strong> o<strong>the</strong>r parts <strong>of</strong> <strong>the</strong> globe.Mammalian remains in<strong>the</strong> Secondary Formations.In <strong>the</strong> oldest Tertiary beds <strong>of</strong> Europe and North America, wehave (even <strong>with</strong> our present imperfect record) a rich and variedmammalian fauna. As compared <strong>with</strong> our living or recent highlyspecialized forms, it may be said to consist <strong>of</strong> generalised types ;but as compared <strong>with</strong> any primeval mammalian type, it must bepronounced highly specialized. Not only are such diversified

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