The geographical distribution of animals, with a study of the relations ...

The geographical distribution of animals, with a study of the relations ... The geographical distribution of animals, with a study of the relations ...

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158 EXTINCT ANIMALS OF AUSTRALIA. [part ii.tion could have occurred in a country like Australia ; but if theocean sank 2,000 feet, the great eastern mountain range mighthave given rise to local glaciers. It is, however, almost certainthat during late Tertiary times Australia must have been muchmore extensive than it is now. This is necessary to allow of thedevelopment of its peculiar and extensive fauna, especially as wesee that that fauna comprised animals rivalling in bulk those ofthe great continents. It is further indicated by the relationswith New Guinea, already alluded to, and by the general characterof the various faunas which compose the Australian region, detailsof which will be found in the succeeding part of this work.The lowering of the ocean during the Glacial period would befavourable to the stillfurther development of the fauna of sucha country ; and it is to the unfavourable conditions produced byits subsequent rising—equivalent to a depression of the land tothe amount of two thousand feet—that we must impute theextinction of so many remarkable groups of animals. It is notimprobable, that the disappearance of the ice and the consequent(apparent) subsidence of the land, might have been rapid ascompared with the rate at which large animals can becomemodified to meet new conditions. Extensive tracts of fertileland might have been submerged, and theconsequent crowdingof large numbers of species and individuals on limited areaswould have led to a struggle for existence in which the lessadapted and less easily modifiable, not the physically weaker,would succumb.There is, however, another cause for the extinction of largerather than small animals whenever an important change ofconditions occurs, which has been suggested to me by a correspondent,1 but which has not, I believe, been adduced by Mr.Darwin or by any other writer on the subject. It is dependenton the fact, that large animals as compared with small ones arealmost invariably slow breeders, and as they also necessarilyexist in much smaller numbers in a given area, they offer farless materials forIn such an extreme case as thatfavourable variations than do smaller animals.1Mr. John Hiekman of Desborongh.of the rabbit and elephant, the

chap, viii.] MESOZOIC MAMMALIA. 159young born each year in the world are probably as some millionsto one ; and it is very easily conceivable that in a thousandyears the former might, under pressure of rapidly changing conditions,become modified into a distinct species, while the latter,not offering enough favourable variations to effect a suitableadaptation, would become extinct. We must also rememberthe extreme specialization of many of the large animals thathave become extinct—a specializationwhich would necessarilyrender modification in any new direction difficult, since the inheritedtendency of variation would probably be to increase thespecialization in the same directionswhich had heretofore beenbeneficial. If to these two causes we add the difficulty ofobtaining sufficient food for such large animals, and perhaps theinjurious effects of changes of climate, we shall not find it difficultto understand how such a vast physical revolution as theGlacial epoch, with its attendant phenomena of elevations andsubsidences, icy winds, and sudden floods by the bursting of lakebarriers, might have led to the total extinction of a vast numberof the most bulky forms of mammalia, while the less bulkywere able to survive, eitherby greater hardiness of constitutionor bv becoming more or less modified. The result is apparent intrie comparatively small or moderate size of the species constitutingthe temperate fauna, in all parts of the globe.It is much to be regretted that no mammalian remains of earlierdate have been found in Australia, as we should then see ifit isreally the case that marsupials have always formed its highesttype of mammalian life. At present its fossil fauna is chieflyinteresting to the zoologist, but throws little light on the pastrelations of this isolated country with other parts of the globe.Mammalian remains inthe Secondary Formations.In the oldest Tertiary beds of Europe and North America, wehave (even with our present imperfect record) a rich and variedmammalian fauna. As compared with our living or recent highlyspecialized forms, it may be said to consist of generalised types ;but as compared with any primeval mammalian type, it must bepronounced highly specialized. Not only are such diversified

158 EXTINCT ANIMALS OF AUSTRALIA. [part ii.tion could have occurred in a country like Australia ; but if <strong>the</strong>ocean sank 2,000 feet, <strong>the</strong> great eastern mountain range mighthave given rise to local glaciers. It is, however, almost certainthat during late Tertiary times Australia must have been muchmore extensive than it is now. This is necessary to allow <strong>of</strong> <strong>the</strong>development <strong>of</strong> its peculiar and extensive fauna, especially as wesee that that fauna comprised <strong>animals</strong> rivalling in bulk those <strong>of</strong><strong>the</strong> great continents. It is fur<strong>the</strong>r indicated by <strong>the</strong> <strong>relations</strong><strong>with</strong> New Guinea, already alluded to, and by <strong>the</strong> general character<strong>of</strong> <strong>the</strong> various faunas which compose <strong>the</strong> Australian region, details<strong>of</strong> which will be found in <strong>the</strong> succeeding part <strong>of</strong> this work.<strong>The</strong> lowering <strong>of</strong> <strong>the</strong> ocean during <strong>the</strong> Glacial period would befavourable to <strong>the</strong> stillfur<strong>the</strong>r development <strong>of</strong> <strong>the</strong> fauna <strong>of</strong> sucha country ; and it is to <strong>the</strong> unfavourable conditions produced byits subsequent rising—equivalent to a depression <strong>of</strong> <strong>the</strong> land to<strong>the</strong> amount <strong>of</strong> two thousand feet—that we must impute <strong>the</strong>extinction <strong>of</strong> so many remarkable groups <strong>of</strong> <strong>animals</strong>. It is notimprobable, that <strong>the</strong> disappearance <strong>of</strong> <strong>the</strong> ice and <strong>the</strong> consequent(apparent) subsidence <strong>of</strong> <strong>the</strong> land, might have been rapid ascompared <strong>with</strong> <strong>the</strong> rate at which large <strong>animals</strong> can becomemodified to meet new conditions. Extensive tracts <strong>of</strong> fertileland might have been submerged, and <strong>the</strong>consequent crowding<strong>of</strong> large numbers <strong>of</strong> species and individuals on limited areaswould have led to a struggle for existence in which <strong>the</strong> lessadapted and less easily modifiable, not <strong>the</strong> physically weaker,would succumb.<strong>The</strong>re is, however, ano<strong>the</strong>r cause for <strong>the</strong> extinction <strong>of</strong> largera<strong>the</strong>r than small <strong>animals</strong> whenever an important change <strong>of</strong>conditions occurs, which has been suggested to me by a correspondent,1 but which has not, I believe, been adduced by Mr.Darwin or by any o<strong>the</strong>r writer on <strong>the</strong> subject. It is dependenton <strong>the</strong> fact, that large <strong>animals</strong> as compared <strong>with</strong> small ones arealmost invariably slow breeders, and as <strong>the</strong>y also necessarilyexist in much smaller numbers in a given area, <strong>the</strong>y <strong>of</strong>fer farless materials forIn such an extreme case as thatfavourable variations than do smaller <strong>animals</strong>.1Mr. John Hiekman <strong>of</strong> Desborongh.<strong>of</strong> <strong>the</strong> rabbit and elephant, <strong>the</strong>

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