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The geographical distribution of animals, with a study of the relations ...

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chap, vii.] MAMMALIA OF THE NEW WORLD. 135also in <strong>the</strong> European Miocene and Upper Eocene formations, andconstitutes a distinct family Hyaenodontidse, allied,according toDr. Leidy, to wolves, cats, hyaenas and weasels. <strong>The</strong> Ursidaeare represented by only one species <strong>of</strong> an extinct genus, Leptarchus,from <strong>the</strong> Pliocene <strong>of</strong> Nebraska. From <strong>the</strong> Pliocene <strong>of</strong>Colorado, Pr<strong>of</strong>. Cope has recently described Tomarctos, as a" short-faced type <strong>of</strong> dog ;" as well as species <strong>of</strong> Cains andMartes.Ungulata.—<strong>The</strong> <strong>animals</strong> belonging to this order being usually<strong>of</strong> large size and accustomed to feed and travel in herds, areliable to wholesale destruction by floods, bogs, precipices, droughtor hunger. It is for <strong>the</strong>se reasons, probably, that <strong>the</strong>ir remainsare almost always more numerous than those <strong>of</strong> o<strong>the</strong>r orders <strong>of</strong>mammalia. In America <strong>the</strong>y are especially abundant ; and <strong>the</strong>number <strong>of</strong> new and intermediate types about whose position<strong>the</strong>re is much difference <strong>of</strong> opinion among Palaeontologists, rendersit very difficult to give a connected summary <strong>of</strong> <strong>the</strong>m <strong>with</strong>any approach to systematic accuracy.Beginning <strong>with</strong> <strong>the</strong> Perissodactyla,or odd-toed ungulates, wefind <strong>the</strong> Equine <strong>animals</strong> remarkably numerous and interesting.<strong>The</strong> true horses <strong>of</strong> <strong>the</strong> genus Equus, so abundant in <strong>the</strong> Post-Pliocene formations, are represented in <strong>the</strong> Pliocene by severalancestral forms.<strong>The</strong> most nearly allied to Equus is Pliohippus,consisting <strong>of</strong> <strong>animals</strong> about <strong>the</strong> size <strong>of</strong> an ass, <strong>with</strong> <strong>the</strong> lateraltoes not externally developed, but <strong>with</strong> some differences <strong>of</strong> dentition.Next come Protohippus and Hipparion, in which <strong>the</strong>lateral toes are developed but are small and functionless. <strong>The</strong>nwe have <strong>the</strong> allied genera, Anchippus, Merychippus, and Hyohippus,related to <strong>the</strong> European Hippo<strong>the</strong>rium, which were all stillsmaller <strong>animals</strong>, Protohippus being only 2| feet high. In <strong>the</strong>older deposits we come to a series <strong>of</strong> forms, still unmistakablyequine, but <strong>with</strong> three or more toes used for locomotion and <strong>with</strong>numerous differentiations in form, proportions, and dentition.<strong>The</strong>se constitute <strong>the</strong> family Anchi<strong>the</strong>ridae. In <strong>the</strong> MioceDe wehave <strong>the</strong> genera Anchi<strong>the</strong>rium (found also in <strong>the</strong> EuropeanMiocene), Miohippus and Mesohippus, all <strong>with</strong> three toes on eachfoot, and about <strong>the</strong> size <strong>of</strong> a sheep or large goat.In <strong>the</strong> Eocene <strong>of</strong>

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