The geographical distribution of animals, with a study of the relations ...
The geographical distribution of animals, with a study of the relations ... The geographical distribution of animals, with a study of the relations ...
126 DISTRIBUTION OF EXTINCT ANIMALS. [part ii.but in some of the species reaching a larger size ; Propalaiotheriumand Pachynolophus of smaller size and having affinitiesfor the other genera named ; and Plagiolopfius, a small, slenderanimal which Professor Huxley thinks may have been a directancestor of the horse. In the Lower Eocene we meet withCoryphodon, much larger than the tapir,and armed with largecanine teeth ; Pliohphus, a generalised type, allied to the tapirand horse ; and Hyracotherium, a small animal from the LowerEocene of England, remotely allied to the tapir.Among the Artiodactyla, or even-toed ungulates, the swineare represented by several extinct genera, of moderate or smalls ize— Acotherium,Chceropotamus, Cebochmrus and Dichobune, allfrom the Upper and the last also from the Middle Eocene ofFrance ; but Eutelodon, from the phosphate of lime deposits islarge. The Dichobune was the most generalised type, presentingthe characters of many of the other genera combined,and was believed by Dr. Falconer to approach the musk-deer.The Cainotherium of the Miocene also occurs here, and an alliedgenus Plesiom&ryx from the same deposits as Euteledon.The Eocene Anoplotheridae were numerous. The Anoplotheriumwas a two-toed, long-tailed Pachyderm, ranging from thesize of a hog to that of an ass ; the allied Eurytherium wasfour-toed ; and there are one or two others of doubtful affinity.All are from the Upper Eocene of France and England.Rodentia.—Eemains referred to the genera Myoxus (dormouse)and Sciurus (squirrel) have been found in the Upper Eoceneof France ; as well as Plesiarctomys, an extinct genus betweenthe marmots and squirrels. The Miocene Theridomys is alsofound here.Marsupials.—The Didclphys (opossum) of Cuvier, now referredto an extinct genus Peratherium, isof France and England.found in the Upper EoceneGeneral Considerations on the Extinct Mammalian Fauna ofEurope.— It is a curious fact that no family, and hardly a genus,of European mammalia occurs in the Pliocene deposits, withoutextending back also into those of Miocene age.There are, how-
;chap, vi.] MAMMALIA OF THE OLD WORLD. 127ever, a few groups which seem to be late developments or recentimportations into the Palaearctic region, as they occur only inPost-Pliocene deposits. The most important of these are thebadger, glutton, elk, reindeer, chamois, goat, and sheep, whichonly occur in caves and other deposits of Post-Pliocene age.Camels only occur in the Post-Pliocene of Siberia (Merycotherium),although a true Camelus of large size appears to have inhabitedsome part of Central Asia in the Upper Miocene period, beingfound in the Siwalik beds. The only exclusively Pliocenegenera in Europe are Ursus, Equits, Hippopotamus, Bos,Elephas,Arvicola, Trogontherium, Arctomys, Hystrix and Lepus ; but ofthese Equus, Hippopotamus, Bos, and Elephas are found in theMiocene deposits of India. Owing, no doubt, in part to thesuperior productiveness of the various Miocene beds, largenumbers of groups appear to have their origin or earliest appearancehere. Such are Insectivora, Felidae, Hyaenidae, Mustelidae,Ursus, Equidae, Tapirus, Ehinocerotidae, Hippopotamidae, Anthracotheridae(extinct), Sus, Camelopardidae, Tragulidae, Cervidae,Bovidse, Elephantidae, and Edentata.Groups which go back to the Eocene period, are, Primatesallied to South American monkeys, as well as some of theLernuridae ; bats of the living genus Vespertilio ; Hyaenodontida?,an ancestral form of Carnivore ; Viverridae ; Canidae (to theUpper Eocene), and the ancestral Arctocyonidae to the LowerEocene ; Hijamarctos, an ancestral type of bears and hyaenasAnchitheridae, ancestral horses, to the Middle Eocene ; Palaeotheridae,comprising numerous generalised forms, ancestors of therhinoceros, horse, and tapir ; Suidoe, with numerous generalisedforms, to the Middle Eocene ;Anoplotheridae and Xiphodontidae,ancestral families of even-toed Ungulates, connecting the ruminantswith the swine ; and lastly, several groups of Eodents,and a Marsupial, in the Upper Eocene. We thus find allthe great types of Mammalia well developed in the earliestportion of the tertiary period ; and the occurrence of Quadrumana, of the highly specialized bats {Vespertilio), of variousforms of Carnivora, and ofUngulates, clearly differentiated intothe odd and even-toed series, associated with such lower forms a&
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;chap, vi.] MAMMALIA OF THE OLD WORLD. 127ever, a few groups which seem to be late developments or recentimportations into <strong>the</strong> Palaearctic region, as <strong>the</strong>y occur only inPost-Pliocene deposits. <strong>The</strong> most important <strong>of</strong> <strong>the</strong>se are <strong>the</strong>badger, glutton, elk, reindeer, chamois, goat, and sheep, whichonly occur in caves and o<strong>the</strong>r deposits <strong>of</strong> Post-Pliocene age.Camels only occur in <strong>the</strong> Post-Pliocene <strong>of</strong> Siberia (Meryco<strong>the</strong>rium),although a true Camelus <strong>of</strong> large size appears to have inhabitedsome part <strong>of</strong> Central Asia in <strong>the</strong> Upper Miocene period, beingfound in <strong>the</strong> Siwalik beds. <strong>The</strong> only exclusively Pliocenegenera in Europe are Ursus, Equits, Hippopotamus, Bos,Elephas,Arvicola, Trogon<strong>the</strong>rium, Arctomys, Hystrix and Lepus ; but <strong>of</strong><strong>the</strong>se Equus, Hippopotamus, Bos, and Elephas are found in <strong>the</strong>Miocene deposits <strong>of</strong> India. Owing, no doubt, in part to <strong>the</strong>superior productiveness <strong>of</strong> <strong>the</strong> various Miocene beds, largenumbers <strong>of</strong> groups appear to have <strong>the</strong>ir origin or earliest appearancehere. Such are Insectivora, Felidae, Hyaenidae, Mustelidae,Ursus, Equidae, Tapirus, Ehinocerotidae, Hippopotamidae, Anthraco<strong>the</strong>ridae(extinct), Sus, Camelopardidae, Tragulidae, Cervidae,Bovidse, Elephantidae, and Edentata.Groups which go back to <strong>the</strong> Eocene period, are, Primatesallied to South American monkeys, as well as some <strong>of</strong> <strong>the</strong>Lernuridae ; bats <strong>of</strong> <strong>the</strong> living genus Vespertilio ; Hyaenodontida?,an ancestral form <strong>of</strong> Carnivore ; Viverridae ; Canidae (to <strong>the</strong>Upper Eocene), and <strong>the</strong> ancestral Arctocyonidae to <strong>the</strong> LowerEocene ; Hijamarctos, an ancestral type <strong>of</strong> bears and hyaenasAnchi<strong>the</strong>ridae, ancestral horses, to <strong>the</strong> Middle Eocene ; Palaeo<strong>the</strong>ridae,comprising numerous generalised forms, ancestors <strong>of</strong> <strong>the</strong>rhinoceros, horse, and tapir ; Suidoe, <strong>with</strong> numerous generalisedforms, to <strong>the</strong> Middle Eocene ;Anoplo<strong>the</strong>ridae and Xiphodontidae,ancestral families <strong>of</strong> even-toed Ungulates, connecting <strong>the</strong> ruminants<strong>with</strong> <strong>the</strong> swine ; and lastly, several groups <strong>of</strong> Eodents,and a Marsupial, in <strong>the</strong> Upper Eocene. We thus find all<strong>the</strong> great types <strong>of</strong> Mammalia well developed in <strong>the</strong> earliestportion <strong>of</strong> <strong>the</strong> tertiary period ; and <strong>the</strong> occurrence <strong>of</strong> Quadrumana, <strong>of</strong> <strong>the</strong> highly specialized bats {Vespertilio), <strong>of</strong> variousforms <strong>of</strong> Carnivora, and <strong>of</strong>Ungulates, clearly differentiated into<strong>the</strong> odd and even-toed series, associated <strong>with</strong> such lower forms a&