The geographical distribution of animals, with a study of the relations ...

The geographical distribution of animals, with a study of the relations ... The geographical distribution of animals, with a study of the relations ...

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72 DISTRIBUTION OF ANIMALS. [part I.so clearly indicated as in some of the other regions, and they areadopted more for convenience than because they are very naturalor strongly marked.The first, or European sub-region, comprises Central andNorthern Europe as far South as the Pyrenees, the Maritimeand Dinaric Alps, the Balkan mountains, the Black Sea, and theCaucasus. On the east the Caspian sea and the Ural mountainsseem the most obvious limit ; but it is doubtful if they form theactual boundary, which is perhaps better marked by the valleyof the Irtish, where a pre-glacial sea almost certainly connectedthe Aral and Caspian seas with the Arctic ocean, and formedan effective barrier which must still, to some extent, influencethe distribution of animals.The next, or Mediterranean sub-region, comprises SouthEurope, North Africa with the extra-tropical portion of theSahara, and Egypt to about the first or second cataracts ; andeastward through Asia Minor, Persia, and Cabul, to the desertsof the Indus.The third, or Siberian sub-region, consists of all north andcentral Asia north of Herat, as far as the eastern limits of thegreat desert plateau of Mongolia, and southward to about theupper limit of trees on the Himalayas.The fourth, or Manchurian sub-region, consists of Japan andNorth China with the lower valley of the Amoor ;and it shouldprobably be extended westward in a narrow strip along theHimalayas, embracing about 1,000 or 2,000 feet of verticaldistance below the upper limit of trees, till it meets an easternextension of the Mediterranean sub-region a little beyond Simla.These extensions are necessary to avoid passing from the Orientalregion, which isessentially tropical, directly to the Siberian subregion,which has an extreme northern character ; whereas theMediterranean and Manchurian sub-regions are more temperatein climate. It will be found that between the upper limit ofmost of the typical Oriental groups and the Thibetan or Siberianfauna, there is a zone in which many forms occur common totemperate China. This is especially the case among the pheasantsand finches.

7;chap, iv.] ZOOLOGICAL REGIONS. 73Ethiopian Region.—The limits of this region have been indicatedby the definition of the Palsearctic region.Besides Africasouth of the tropic of Cancer, and its islands, it comprises thesouthern half of Arabia.This region has been said to be identical in the main charactersof its mammalian fauna with the Oriental region, and hastherefore been united with it by Mr. A. Murray. Most importantdifferences have however been overlooked, as the followingsummary of the peculiarities of the Ethiopian region will, Ithink, show.It possesses 22 peculiar families of vertebrates ; 90 peculiargenera of mammalia, being two-thirds of its whole numberand 179 peculiar genera of birds, being three-fifths of all itpossesses. It is further characterized by the absence of severalfamilies and genera which range over the whole northernhemisphere, details of which will be found in the chaptertreating of the region. There are, it is true, many pointsof resemblance, not to be wondered at between two tropicalregionsin the same hemisphere, and which have evidently beenat one time more nearly connected, both by intervening landsand by a different condition of the lands that even now connectthem.remarkable ;But these resemblances only render the differences moresince they show that there has been an ancient andlong-continued separation of the two regions, developing a distinctfauna in each, and establishing marked specialitieswhichthe temporary intercommunication and immigration has notsufficed to remove. The entire absence of such wide-spreadgroups as bears and deer, from a country many parts of whichare well adapted to them, and in close proximity to regionswhere they abound, would alone mark out the Ethiopian regionas one of the primary divisions of the earth, even if it possesseda less number than it actually does of peculiar family andgenericgroups.Sub-divisions of the Ethiopian Region.—The African continentsouth of the tropic of Cancer is more homogeneous in itsprominent and superficial zoological features than most of theother regions, but there are nevertheless important and deep-Vol. I.—

72 DISTRIBUTION OF ANIMALS. [part I.so clearly indicated as in some <strong>of</strong> <strong>the</strong> o<strong>the</strong>r regions, and <strong>the</strong>y areadopted more for convenience than because <strong>the</strong>y are very naturalor strongly marked.<strong>The</strong> first, or European sub-region, comprises Central andNor<strong>the</strong>rn Europe as far South as <strong>the</strong> Pyrenees, <strong>the</strong> Maritimeand Dinaric Alps, <strong>the</strong> Balkan mountains, <strong>the</strong> Black Sea, and <strong>the</strong>Caucasus. On <strong>the</strong> east <strong>the</strong> Caspian sea and <strong>the</strong> Ural mountainsseem <strong>the</strong> most obvious limit ; but it is doubtful if <strong>the</strong>y form <strong>the</strong>actual boundary, which is perhaps better marked by <strong>the</strong> valley<strong>of</strong> <strong>the</strong> Irtish, where a pre-glacial sea almost certainly connected<strong>the</strong> Aral and Caspian seas <strong>with</strong> <strong>the</strong> Arctic ocean, and formedan effective barrier which must still, to some extent, influence<strong>the</strong> <strong>distribution</strong> <strong>of</strong> <strong>animals</strong>.<strong>The</strong> next, or Mediterranean sub-region, comprises SouthEurope, North Africa <strong>with</strong> <strong>the</strong> extra-tropical portion <strong>of</strong> <strong>the</strong>Sahara, and Egypt to about <strong>the</strong> first or second cataracts ; andeastward through Asia Minor, Persia, and Cabul, to <strong>the</strong> deserts<strong>of</strong> <strong>the</strong> Indus.<strong>The</strong> third, or Siberian sub-region, consists <strong>of</strong> all north andcentral Asia north <strong>of</strong> Herat, as far as <strong>the</strong> eastern limits <strong>of</strong> <strong>the</strong>great desert plateau <strong>of</strong> Mongolia, and southward to about <strong>the</strong>upper limit <strong>of</strong> trees on <strong>the</strong> Himalayas.<strong>The</strong> fourth, or Manchurian sub-region, consists <strong>of</strong> Japan andNorth China <strong>with</strong> <strong>the</strong> lower valley <strong>of</strong> <strong>the</strong> Amoor ;and it shouldprobably be extended westward in a narrow strip along <strong>the</strong>Himalayas, embracing about 1,000 or 2,000 feet <strong>of</strong> verticaldistance below <strong>the</strong> upper limit <strong>of</strong> trees, till it meets an easternextension <strong>of</strong> <strong>the</strong> Mediterranean sub-region a little beyond Simla.<strong>The</strong>se extensions are necessary to avoid passing from <strong>the</strong> Orientalregion, which isessentially tropical, directly to <strong>the</strong> Siberian subregion,which has an extreme nor<strong>the</strong>rn character ; whereas <strong>the</strong>Mediterranean and Manchurian sub-regions are more temperatein climate. It will be found that between <strong>the</strong> upper limit <strong>of</strong>most <strong>of</strong> <strong>the</strong> typical Oriental groups and <strong>the</strong> Thibetan or Siberianfauna, <strong>the</strong>re is a zone in which many forms occur common totemperate China. This is especially <strong>the</strong> case among <strong>the</strong> pheasantsand finches.

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