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LILIACEAE - China

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Mixed forests; 1600–1800 m. SE Yunnan (Malipo Xian).64. Smilax cocculoides Warburg in Diels, Bot. Jahrb. Syst. 29:257. 1900.银 叶 菝 葜 yin ye ba qiaSmilax polycolea Warburg var. acuminata Warburg.Shrubs subscandent, unarmed. Stem suberect, branched,0.5–2 m. Petiole usually curved, 5–10 mm, transversely wrinkledon proximal 1/2, basally narrowly winged; wings sometimesauriculate apically; abscission zone at middle; tendrilsabsent. Leaf blade abaxially grayish green, ovate, elliptic-ovate,or ovate-lanceolate, abaxially slightly shiny. Male inflorescenceof 1(or 2) umbels, basally prophyllate; peduncle 1–2 cm, articulate2–5 mm from base; umbels 3–15-flowered, base scarcelythickened. Male flowers: tepals yellowish green, outer ones2.5–3.5 × 1–1.5 mm; stamens very short, ca. 0.7 mm. Femaleflowers unknown. Berries blackish blue, globose, ca. 8 mm indiam. Fl. Feb–Apr, fr. Nov.Forests, thickets; 500–1900 m. Guangdong, Guangxi, Guizhou,Hubei, Hunan, Sichuan, Yunnan.65. Smilax pinfaensis H. Léveillé & Vaniot in H. Léveillé,Mem. Pontif. Accad. Romana Nuovi Lincei 23: 355. 1905.平 伐 菝 葜 ping fa ba qiaShrubs suberect, unarmed. Rhizome stolonlike, 2–3 mm indiam. Stem branched, smooth. Petiole 4–6 cm, basally narrowlywinged; abscission zone apical; tendrils absent. Leaf blade lanceolateto oblong-lanceolate, 8–16 × 1.5–4 cm, leathery, 3-veined, with midvein concave adaxially, base subrounded, apexacuminate. Male inflorescence of 1 umbel, basally prophyllate;peduncle 8–14(–20) cm, proximally articulate; umbel 5- or 6-flowered, base scarcely thickened; bracteoles small; pedicels 5–9 mm. Male flowers: tepals spreading, ovate or ovate-elliptic,1.5–1.8 × ca. 1 mm; stamens very short, 0.3–0.4 mm; anthersshorter than filaments. Female flowers unknown. Fl. Mar.Forests. SC Guizhou (Guiding Xian).66. Smilax basilata F. T. Wang & Tang, Fl. Reipubl. PopularisSin. 15: 254. 1978.少 花 菝 葜 shao hua ba qiaVines climbing. Stem branched, to 2 m, woody; branches2–4-angled, sometimes sparsely prickly. Petiole 2.5–5 mm, narrowlywinged for 1/2–2/3 its length; abscission zone distal; tendrilssometimes present. Leaf blade lanceolate, ovatelanceolate,or oblong-lanceolate, 3–7 × 1–2(–2.5) cm, leatheryor thickly papery. Male inflorescence of 1 umbel, basallyprophyllate; peduncle 1–3 mm, much shorter than petiole,proximally articulate; umbels 2- or 3-flowered, base scarcelythickened. Male flowers: tepals greenish, outer ones ca. 2 × 1mm, inner ones ca. 1 × 0.6 mm; stamens ca. 1/2 as long astepals; anthers suborbicular. Female flowers unknown. Berriesdark purple, globose, ca. 1 cm in diam. Fl. Mar–Apr, fr. Nov.Forests, grassy slopes; 1200–2000 m. Guangxi, SE Yunnan.The recently described Smilax guiyangensis C. X. Fu & C. D.Shen (Acta Phytotax. Sin. 35: 70. 1997), from C Guizhou (Guiyang),appears to be a distinct species closely related to S. basilata, fromwhich it differs in having white-marked leaf blades and rather longpeduncles conspicuously longer than the petioles.67. Smilax fooningensis F. T. Wang & Tang, Fl. Reipubl. PopularisSin. 15: 255. 1978.富 宁 菝 葜 fu ning ba qiaVines climbing. Stem branched, woody; branches denselyverruculose. Petiole 1–2 cm, wingless; abscission zone betweenmiddle and distal part; tendrils absent. Leaf blade ovate,7.5–15.5 × 4–8 cm, leathery, base subtruncate or broadly cuneate,apex acuminate. Male inflorescence of 1 umbel, basallyprophyllate; peduncle 3–5 cm, proximally articulate; umbeldensely many flowered, base thickened, oblong-ellipsoid, 4–5 ×1.5–2 mm. Male flowers: tepals yellowish green, outer onesoblong, 2–2.5 × ca. 0.8 mm, inner ones ca. 2 × 0.4 mm; stamensca. 1/2 as long as tepals; anthers oblong, ca. 0.5 mm. Femaleflowers and fruit unknown. Fl. May.Forests; ca. 600 m. SE Yunnan (Funing Xian).68. Smilax quadrataA. de Candolle in A. de Candolle & C. deCandolle, Monogr. Phan. 1: 183. 1878.方 枝 菝 葜 fang zhi ba qiaVines climbing. Stem branched, woody; branches stronglyzigzagged, obtusely 4-angled, subsquare in cross section, sometimessparsely prickly. Petiole 1–1.7 cm, narrowly winged for1/2–2/3 its length; abscission zone distal; tendrils usually present.Leaf blade oblong to ovate-oblong, 7–11 × 2.5–4.5 cm,thickly papery or thinly leathery, base subtruncate or rounded,apex mucronate. Male inflorescence a raceme of 2–4 umbels,2.5–4 cm, basally prophyllate; umbels 20–30-flowered, baseslightly thickened. Male flowers: tepals 4–5 × ca. 1.5 mm, innerones ca. 5 × 0.5 mm; stamens 4–5 mm; anthers linear, ca. 1.2mm. Female flowers and fruit unknown. Fl. Oct.Forests, shaded places along valleys; 1900–2000 m. ?Xizang, SEand WC Yunnan (Jingdong Xian, Yanshan Xian) [India, Myanmar].69. Smilax lunglingensis F. T. Wang & Tang, Bull. Fan Mem.Inst. Biol. 7: 86. 1936.马 钱 叶 菝 葜 ma qian ye ba qiaSmilax siderophylla Handel-Mazzetti.Vines climbing. Stem branched, subterete, 1–3 m or more,woody;branches usually sparsely prickly. Petiole 2–3 cm, winglessor rarely narrowly winged for ca. 1/2 its length; abscissionzone distal; tendrils usually present. Leaf blade elliptic, ovateelliptic,or orbicular, 5–11 × 3–9 cm, thickly leathery, main veins3, strongly convex abaxially, concave adaxially. Inflorescence araceme of 2–4 umbels, 3–5 cm, basally prophyllate; umbels ofboth sexes 15–25-flowered, base subglobose. Male flowers:tepals yellow, outer ones ca. 6 × 2 mm, inner ones much narrower;stamens ca. 6 mm. Female flowers: tepals smaller thanmale ones; staminodes 3. Berries black, globose, 5–6 mm indiam. Fl. Mar–Apr, fr. Aug–Oct.Forests, thickets, shaded and damp places on slopes; 1800–2700m. Yunnan.

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