Plants stoloniferous; stolons with spaced nodes. Leavesbasal, tufted; petiole 2.5–10 cm; leaf blade abaxially glaucous,narrowly oblong to suboblanceolate, 5–12 × 0.5–1.3 cm, 5–7-veined, base attenuate, apex subrounded or obtuse. Scape 7–11cm. Inflorescence a raceme, 1–3(or 4)-flowered; bracts ovate,ca. 7 mm, margin membranous. Flowers solitary; pedicel 5–8 mm, articulate near apex. Tepals scarcely spreading, white,tinged purplish, suboblong, ca. 1.2 cm × 3–4.5 mm. Filamentsca. 2 mm; anthers ca. 7 mm. Style ca. 1 cm, slender. Seeds deepblue at maturity, ellipsoid, ca. 8 mm in diam. Fl. May–Jun, fr.Jul–Aug. 2n = 36*.Forests, sparse forests, hillsides along streams; 1000–1600 m.Guangdong, Guangxi, NE Guizhou (Fanjing Shan), Hubei, Hunan, Sichuan.31. Ophiopogon pingbienensis F. T. Wang & L. K. Dai in F.T. Wang & Tang, Fl. Reipubl. Popularis Sin. 15: 251. 1978.屏 边 沿 阶 草 ping bian yan jie caoStem very short, indistinct. Leaves basal, tufted; petiole 4–14 cm; leaf blade abaxially slightly glaucous, narrowly ellipticoblong,5.5–9 × 1.5–2 cm, base attenuate, apex acute. Scape 6–10 cm. Inflorescence a raceme, several flowered; bracts ovate toovate-lanceolate, 6–8 mm. Flowers solitary; pedicel 4–6 mm,articulate at middle. Tepals oblong-lanceolate to narrowly oblong,8–11 × 2.5–3.5 mm. Filaments ca. 1 mm; anthers ca. 6mm. Style ca. 7 mm, slender. Fl. May. 2n = 36*.Dense forests; 1800–2000 m. SE Yunnan (Pingbian Miao ZuZizhixian).32. Ophiopogon tonkinensis L. Rodriguez, Bull. Soc. Bot.France 75: 998. 1928.多 花 沿 阶 草 duo hua yan jie caoRhizome short, with densely white-hairy roots. Leavesbasal, tufted; petiole (6–)10–20(–28) cm; leaf blade oblanceolate-oblong,15–25 × 2.5–3.5(–4.2) cm, thickly leathery, apexacute or mucronate. Scape 15–24 cm. Inflorescence a reducedpanicle, 9–12 cm, 15–35-flowered; bracts ovate to lanceolate,basal one 6–9 mm. Flowers usually in clusters of 2–4; pedicel3–5 mm, articulate at middle. Tepals purplish, ovate to oblong,ca. 4 × 2 mm. Filaments ca. 1 mm; anthers ca. 3 mm. Style 3–4mm, slender. Seeds ellipsoid to globose, ca. 9 mm in diam. Fl.Sep, fr. Oct–Nov. 2n = 36*.Dense forests, scrub forests, thickets, open grassy slopes; 1000–1600 m. W Guangxi, SE Yunnan [Vietnam].33. Ophiopogon pseudotonkinensis D. Fang, J. Trop. Subtrop.Bot. 6: 100. 1998.拟 多 花 沿 阶 草 ni duo hua yan jie caoRhizome ca. 2.5 × 1 cm, robust. Roots ca. 1.5 mm thick.Leaves basal, tufted; petiole 4–8 cm; leaf blade usually narrowlyelliptic-oblong, rarely linear, asymmetric, 13.5–27 × 1–2.2 cm, papery, 9–16-veined, base attenuate, apex ± obtuse.Scape ca. 5.5 cm, compressed. Inflorescence a reduced panicle,ca. 7 cm, ca. 30-flowered; bracts narrowly ovate, membranous,basal one ca. 7 mm, base ca. 3 mm wide. Flowers in clusters of2 or 3; pedicel ca. 1 cm, articulate at middle. Tepals reflexeddistally, white, ovate, ca. 5 × 3.5–4 mm, apex obtuse. Filamentsvery short; anthers ovate-oblong, ca. 3 mm, connate, apex subtruncate.Style ca. 5 mm. Fl. Sep.Limestone hillsides. NW Guangxi (Nandan Xian).34. Ophiopogon marmoratus Pierre ex L. Rodriguez, Bull.Soc. Bot. France 75: 997. 1928.丽 叶 沿 阶 草 li ye yan jie caoRoots slender, soft. Leaves basal, tufted; petiole 8–12 cm;leaf blade narrowly oblong, 13–18 × 1.8–2.5 cm, distinctly ca.7-veined, base attenuate, apex acute to shortly acuminate. Scape15–30 cm. Inflorescence a reduced panicle,10–13 cm, laxly 15–20-flowered; bracts ovate to broadly so, membranous, basal oneca. 1 cm. Flowers solitary or in clusters of 2 or 3; pedicel ca. 8mm, articulate proximally. Tepals white, oblong-lanceolate, ca.8 × 1.5–2.5 mm. Filaments very short; anthers ca. 4 mm. Styleca. 8 mm. Fl. Aug. 2n = 36*.Dense forests along valleys. SW Guangxi, SE Yunnan [Cambodia,Laos, Thailand, Vietnam].35. Ophiopogon xylorrhizus F. T. Wang & L. K. Dai in F. T.Wang & Tang, Fl. Reipubl. Popularis Sin. 15: 252. 1978.木 根 沿 阶 草 mu gen yan jie caoRoots somewhat stiltlike, straight, 3–5 mm thick, stiff, ±woody, not hairy. Leaves basal, tufted; petiole 7–9 cm; leafblade suboblong, 11–16 × 3.2–4.2 cm, base attenuate, apexshortly acuminate. Scape ca. 10 cm. Inflorescence a reducedpanicle, ca. 6 cm, more than 20-flowered; bracts ovate, 4–5mm. Flowers solitary or in clusters of 2 or 3; pedicel ca. 3 mm,articulate at middle. Tepals bluish, ovate-lanceolate to lanceolate,ca. 5 × 2 mm. Filaments ca. 1 mm; anthers 2–2.5 mm.Style ca. 4 mm. Fl. Jun. 2n = 36*.Dense forests, rainforests, thickets, moist and shady places;600–1200 m. S Yunnan.36. Ophiopogon zingiberaceus F. T. Wang & L. K. Dai in F.T. Wang & Tang, Fl. Reipubl. Popularis Sin. 15: 252. 1978.姜 状 沿 阶 草 jiang zhuang yan jie caoRhizome gingerlike, ca. 3 cm thick, fleshy. Leaves basal,tufted, sessile, grasslike, 15–30 cm × 4–6 mm, 5–9-veined.Scape ca. 18 cm. Inflorescence a raceme, ca. 3 cm, more than10-flowered; bracts linear, basal one ca. 7 mm. Flowerssolitary; pedicel ca. 2.5 mm, articulate near middle. Tepalsdeltoid-ovate, ca. 4 mm (in slightly immature flowers).Filaments very short; anthers ca. 3 mm. Style ca. 3 mm. Fl.May–Jun. 2n = 36*.Mixed forests, bamboo forests, moist and shady places; 1400–3000 m. S Sichuan, N and SE Yunnan.37. Ophiopogon megalanthus F. T. Wang & L. K. Dai in F. T.Wang & Tang, Fl. Reipubl. Popularis Sin. 15: 253. 1978.大 花 沿 阶 草 da hua yan jie caoPlants basally ± purple-brown. Rhizome tuberous, thick.Roots densely yellow-brown tomentose when young. Leavesbasal, tufted, sessile, grasslike, 25–60 × 0.8–1.5 cm, marginmembranous at base, serrulate distally; leaf tufts surrounded by
many linear-lanceolate sheaths. Scape 30–35 cm. Inflorescencea reduced panicle, 7–10 cm, 10–25-flowered; bracts lanceolate,slightly falcate, broadly membranous at margin, basal one 1.5–4 cm. Flowers in clusters of 2–4; pedicel 1–1.5 cm, articulateproximally. Tepals purplish red, 9–10 mm, outer ones ovate,inner ones lanceolate. Filaments very short; anthers ca. 5 mm.Style ca. 8 mm. Fl. Jul. 2n = 36*.Dense forests, scrub forests, thickets along streams; 1100–2800m. S Yunnan.38. Ophiopogon revolutus F. T. Wang & L. K. Dai in F. T.Wang & Tang, Fl. Reipubl. Popularis Sin. 15: 253. 1978.卷 瓣 沿 阶 草 juan ban yan jie caoRoots slender, white hairy, sometimes with small,tuberous part near tip. Leaves basal, tufted, indistinctlypetiolate, abaxially glaucous, grasslike, 25–50 × 0.8–1.2 cm,base attenuate, margin serrulate; leaf tufts surrounded bybroadly lanceolate sheaths. Scape 20–40 cm. Inflorescence araceme or a reduced panicle, 8–23 cm, 10–24-flowered; bractssubulate, basal one 16–28 mm. Flowers solitary or sometimespaired; pedicel 7–9 mm, articulate near base. Tepals revolute,white or purplish, linear, ca. 9 × 1.5 mm. Filaments veryshort; anthers ca. 7 mm, slightly connate or free. Style ca. 8mm, slender. Seeds ellipsoid, ca. 9 × 5 mm. Fl. and fr. Sep–Oct. 2n = 36*.Rainforests, evergreen broad-leaved forests, dense or sparse forests;500–1900 m. S Yunnan [Thailand].Tanaka (in J. Jap. Bot. 74: 321–328. 1999) reduced Ophiopogonrevolutus to the synonymy of O. griffithii. However, one of us (Tamura)prefers to maintain the former species and regard the latter as a synonymof the very variableO. intermedius.39. Ophiopogon corifolius F. T. Wang & L. K. Dai in F. T.Wang & Tang, Fl. Reipubl. Popularis Sin. 15: 353. 1978.厚 叶 沿 阶 草 hou ye yan jie caoRoots slender, soft, white hairy. Leaves basal, tufted, indistinctlypetiolate,abaxially glaucous and whitestreaked, grasslike,38–60 × 1–1.5 cm, leathery, rigid, base attenuate, marginslightly recurved; leaf tufts ± surrounded by membranoussheaths. Scape ca. 22 cm. Inflorescence a reduced panicle, ca.12 cm, many flowered; bracts purple-brown, subovate, basalone ca. 6 mm. Flowers usually in clusters of 2–4; pedicel 1–1.2cm, articulate distally. Tepals purple, ca. 9 mm, outer ones linear,inner ones lanceolate. Filaments ca. 1 mm; anthers ca. 8mm. Style ca. 9 mm, slender. Seeds ellipsoid, ca. 10 × 8 mm.Fl. Apr–May, fr. Jul–Aug.Dense forests; 1200–1400 m. Guangxi, SW Guizhou.40. Ophiopogon sparsiflorus F. T. Wang & L. K. Dai in F. T.Wang & Tang, Fl. Reipubl. Popularis Sin. 15: 253. 1978.疏 花 沿 阶 草 shu hua yan jie caoRoots slender, soft, densely white hairy. Leaves basal, tufted,indistinctly petiolate, abaxially glaucous, grasslike, 15–40cm × 4–7(–9) mm, 5–9-veined, base attenuate; leaf tufts surroundedby membranous sheaths. Scape 13–28 cm. Inflorescencea raceme, 6–8 cm, several to 15-flowered; bracts lanceolate,basal one 7–8 mm. Flowers solitary; pedicel ca. 1 cm,articulate near middle. Tepals purplish, narrowly lanceolate, ca.10 × 2.5–3 mm. Filaments ca. 2 mm; anthers ca. 7 mm, initiallyconnate, later free. Style ca. 8 mm, slender. Fl. May.Forests, moist and shady places along streams; 800–1400 m.SW Guangdong, Guangxi.41. Ophiopogon intermedius D. Don, Prodr. Fl. Nepal. 48.1825.间 型 沿 阶 草 jian xing yan jie caoFlueggea dubia Kunth; F. griffithii Baker; F. jacquemontianaKunth; F. japonica (Linnaeus f.) Richard var. intermedia(D. Don) Schultes; F. wallichiana Kunth; Mondo intermedium(D. Don) L. H. Bailey; M. japonicum (Linnaeus f.) Farwell var.griffithii (Baker) Farwell; M. japonicum var. intermedium (D.Don) Farwell; M. japonicum var. wallichianum (Kunth) Farwell;M. scabrum Ohwi; M. wallichianum (Kunth) L. H.Bailey; Ophiopogon aciformis F. T. Wang & Tang ex H. Li &Y. P. Yang; O. compressus Y. Wan & C. C. Huang; O.griffithii (Baker) J. D. Hooker; O. japonicus (Linnaeus f.) KerGawler var. intermedius (D. Don) Maximowicz; O.longibracteatus H. Li & Y. P. Yang; O. longipedicellatus Y.Wan & C. C. Huang; O. wallichianus (Kunth) J. D. Hooker; O.xiaokuai Z. Y. Zhu.Roots sometimes with fleshy, tuberous part near tip. Rhizomeshort, thick. Leaves basal, tufted, sessile, grasslike, 15–55(–70) cm × 3–15 mm, 5–9-veined, base attenuate, margin usuallyserrulate; leaf tufts usually surrounded by brownish, membranoussheaths. Scape (15–)20–50 cm, sometimes slightlyflattened and narrowly 2-winged. Inflorescence a reduced panicle,(2.5–)5–16 cm, many flowered; bracts subulate, linear, orlanceolate, basal one 1.5–4 cm. Flowers in clusters of 2 or 3 orsometimes solitary; pedicel 4–7 mm, articulate proximally or atmiddle. Tepals white or purplish, oblong, ovate, or ovate-lanceolate,4–7 × 2–2.5 mm. Filaments very short; anthers 2–4mm. Style 3–4 mm. Seeds ellipsoid. Fl. May–Aug, fr. Aug–Oct.2n = 36*, 54, 68, 72*, 108*, 112.Evergreen broad-leaved forests, mixed forests, bamboo forests,scrub, moist and shady places along streams, grassy slopes; 700–3000m. Anhui, Guangdong, Guangxi, Guizhou, Hainan, Henan, Hubei, Hunan,Shaanxi, Sichuan, Taiwan, Xizang, Yunnan [Bangladesh, Bhutan,India, Myanmar, Nepal, Sikkim, Sri Lanka, Thailand, Vietnam].Ophiopogon intermedius is a very variable species. Ophiopogongriffithii, reported by Yang and Li (Acta Bot. Yunnan., Suppl. 3: 82–83. 1990) from S Yunnan, seems to be conspecific with O. intermedius.42. Ophiopogon reversus C. C. Huang, Fl. Hainan. 4: 534.1977.高 节 沿 阶 草 gao jie yan jie caoRhizome short. Leaves basal, tufted, sessile, long linear,30–50 cm × 3–8 mm, (5–)7–9-veined, base attenuate, marginserrulate distally. Scape 18–25 cm, slightly flattened and minutelywinged. Inflorescence a reduced panicle, 5–7 cm, manyflowered; bracts deltoid to ovate, basal one to 2 cm. Flowers inclusters of 2 or 3 or solitary; pedicel curved, 5–10 mm, artic-
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34b. Stem very short; leaves basal
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simple, erect, usually with a few s
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ciflorus and S. spicatus Klotzsch);
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pendulous .........................
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ase, persistent. Stamens 6, free or
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Helonias umbellata (Baker) N. Tanak
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6-11 cm, adaxially sometimes with p
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小 重 楼 xiao chong louPlants 7-
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14b. Paris thibetica var. apetala H
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slightly convex connective apically
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when dried, not wrinkled adaxially
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80b. Flowers much smaller, tepals 2
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8. Smilax davidiana A. de Candolle
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present. Leaf blade abaxially sligh
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This species was misidentified in F
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slightly shorter than petiole; umbe
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马 甲 菝 葜 ma jia ba qiaVines
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Vines climbing, usually unarmed. St
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70. Smilax bauhinioides Kunth, Enum
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4a. Male umbel 5-9-flowered; filame
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persistent tepals. Seeds many, glob
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Imag. Descr. Fl. Russ. 8. 1844; Gag
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ca. 2/3 as wide as inner ones.6a. I
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1a. Bracts 2-4, much smaller than l
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oblong to lanceolate, 2-2.5 cm × 6
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12b. Leaves 0.5-1 cm wide; inflores
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Yunnan.The bulbs are used medicinal
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甘 肃 贝 母 gan su bei muFritil
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ather long, slender; stigma 3-lobed
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2b. Flowers not campanulate, tepals
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etween apical whorl and basal bract
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Lilium nanum Klotzsch var. brevisty
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2.5-3 cm × 5-10 mm. Stem 20-30 cm,
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5.5-6 cm × 8-10 mm, margin revolut
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woolly hairs ......................
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3-5-veined, margin papillose. Flowe
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3a. Inner tepals neither spotted no
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3a. Stems subglabrous.4a. Flowers p
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Tepals obliquely outward spreading,
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12a. Tepals white to cream, spotted
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latipetalum Collett & Hemsley; D. p
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only slightly exserted from periant
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laysia, Myanmar, Nepal, Philippines
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3b. Perianth tube longer, to 4 cm,r
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About 660 species: N hemisphere, ma
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44a. Scape 10-15(-20) cm; perianth
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85b. Perianth segments 3-6 mm; fila
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125b. Inner filaments entire at bas
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Leaf blade without white main veins
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irregularly 2-lobed. Filaments subu
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Scrub, moist places; 4500-5000 m. S
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Forests, slopes, pastures; 1600-330
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(40-)50-80 cm, terete, covered with
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2/3-5/6 as long as perianth segment
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Filaments shorter than perianth seg
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umbel, covered with leaf sheaths on
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shores; 1900-3700 m. Xinjiang [Afgh
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昆 仑 韭 kun lun jiuBulbs cluste
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Bulb solitary or clustered, applana
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北 葱 bei congBulbs usually clust
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112. Allium songpanicum J. M. Xu in
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Style exserted. Fl. and fr. May-Jul
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Forests on shady slopes, sands, san
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stigma small. Fruit a capsule, glob
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90: 27. 1931.紫 萼 zi eBryocles v
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Tepals white, linear, membranous; o
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