ase, persistent. Stamens 6, free or basally adnate to perianth, slightly shorter than perianth; filaments filiform or widened in proximalpart; anthers small, suborbicular to reniform, basifixed, extrorse, with confluent locules, dehiscing transversely by valves. Ovarysuperior or semi-inferior, 3-loculed; ovules many. Styles 3, slender. Capsule ovoid to conical, 3-lobed, ventricidal. Seeds many, oblongor oblanceolate, narrowly winged or angular.About ten species: N Arctic to temperate regions, especially in North America; one species in <strong>China</strong>.1. Zigadenus sibiricus (Linnaeus) A. Gray, Ann. Lyceum Nat.Hist. New York 4: 112. 1857.棋 盘 花 qi pan huaMelanthium sibiricum Linnaeus, Sp. Pl. 1: 339. 1753;Anticlea sibirica (Linnaeus) Kunth.Bulb cylindric, rarely to ovoid, slender, with blackishbrown coat. Leaves (10–)13–35 cm × 2–8(–10) mm, glabrous,base tapering to sheath, apex obtuse. Scape (10–)30–50 cm,usually with 1 or 2 slightly smaller leaves in proximal part. Inflorescenceusually racemose, less often 1- or 2-branchedand paniculate, laxly 4–12-flowered; bracts linear-lanceolateto ovate. Pedicel 0.7–2 cm. Flowers erect. Tepals greenishwhite, obovate-oblong or oblong, 6–9 × 2.2–2.6(–4) mm,spreading at anthesis, each with a large, yellowish green,obcordate gland nearly at base. Filaments glabrous, graduallywidened toward base; anthers reniform. Ovary semi-inferior,conical, ca. 4 mm. Styles recurved, extending beyond perianthafter anthesis. Capsule conical, ca. 1.5 cm. Seeds suboblong,ca. 5 mm. Fl. Jul, fr. Aug–Sep. 2n = 32.Forests, damp places in thickets, grassy slopes; near sea level to2600 m. Hebei, Heilongjiang, Hubei, Jilin, Liaoning, Nei Mongol,Shanxi, Sichuan [Japan (Rebun and Rishiri Islands), Korea, Mongolia,Russia].6. YPSILANDRA Franchet, Nouv. Arch. Mus. Hist. Nat., sér. 2, 10: 93. 1887.丫 蕊 花 属 ya rui hua shuChen Xinqi ( 陈 心 启 Chen Sing-chi); Minoru N. TamuraHerbs perennial, with a short, thickened, slightly fleshy rhizome, glabrous. Leaves basal, rosulate, linear to lanceolate or oblanceolate,or spatulate, basally gradually narrowed to a petiole. Scape arising from axils of lateral leaves, erect, simple, with several tomany scaly leaves. Inflorescence a terminal raceme, 2–30-flowered; bract absent. Flowers bisexual, usually nodding at anthesis,ascending in fruit, spreading funnelform. Tepals 6, free, with a nectary gland basally on adaxial side, persistent. Stamens 6, ratherlong, free from tepals, inner ones basally adnate to ovary, outer ones free; anthers usually reniform, basifixed, with confluent locules.Ovary superior, 3-lobed, 3-loculed; ovules many per locule. Style 1, very short to long; stigma capitate to 3-cleft. Fruit a capsule,trigonous, 3-lobed apically, loculicidal. Seeds numerous, narrowly fusiform, both ends caudate.Five species: Bhutan, <strong>China</strong>, Myanmar, Nepal; five species (three endemic) in <strong>China</strong>.1a. Stamens and style not extending beyond tepals at anthesis; stigma shortly or deeply 3-lobed (lobes 0.3–0.8 mm).2a. Stigma deeply 3-lobed (lobes ca. 0.8 mm); style 1–1.5 mm; tepals 4–5 mm, spatulate or oblanceolate toelliptic, shorter than capsule; flowers 5–17 per scape ....................................................................................... 4. Y. yunnanensis2b. Stigma shortly to moderately 3-lobed (lobes 0.3–0.5 mm); style 2.5–6 mm; tepals 7–12 mm, linear to linearlanceolate,longer than capsule; flowers 2–6 per scape .............................................................................................. 5. Y. alpina1b. Stamens and style extending beyond tepals at anthesis; stigma capitate, slightly or scarcely 3-lobed.3a. Scape densely covered with many imbricate, scaly leaves; pedicel shorter than tepals ..................................... 2. Y. kansuensis3b. Scape laxly covered with several scaly leaves; pedicel equaling tepals.4a. Either pedicel or tepals 6–10 mm; stamens and style obviously extending beyond tepals at anthesis; ovarylobes 1/3–2/5 as long as ovary .......................................................................................................................... 1. Y. thibetica4b. Either pedicel or tepals 4–6 mm; stamens and style slightly extending beyond tepals at anthesis; ovarylobes 1/5–1/4 as long as ovary ........................................................................................................................ 3. Y. cavaleriei1. Ypsilandra thibetica Franchet, Nouv. Arch. Mus. Hist. Nat.,sér. 2, 10: 94. 1887.丫 蕊 花 ya rui huaHelonias thibetica (Franchet) N. Tanaka; Ypsilandra thibeticavar. angustifolia F. T. Wang & Tang.Rhizome 1–5 × ca. 1 cm. Leaves usually more than 10,oblanceolate, basally gradually narrowed to a petiole; petiole 3–15 cm × 3–5 mm; leaf blade 4–14 × 1–4.8 cm, apex acuminate.Scape 7–50 cm, usually longer than basal leaves, laxly coveredwith several scaly leaves. Raceme 5–30-flowered. Pedicel 6–10mm, nearly as long as tepals. Tepals white, pink, or purple,spatulate-oblanceolate, 6–10 × 1.5–2.5 mm, 3–5-veined. Stamens1–1.8 cm, obviously extending beyond tepals at anthesis.Ovary deeply 3-lobed apically; lobes 1/3–2/5 as long as ovary.Style 1.6–2 cm, slightly longer than stamens; stigma capitate,slightly or scarcely 3-lobed. Capsule broadly ovoid, 1/2–2/3 aslong as persistent tepals. Seeds 4–5 mm. Fl. Mar–Apr, fr. May–Jun. 2n = 34*.Forests, moist places on hillsides, shady slopes along valleys;1300–2900 m. NE Guangxi, S Hunan, Sichuan.2. Ypsilandra kansuensis R. N. Zhao & Z. X. Peng, Acta Bot.
Bor.-Occid. Sin. 7: 57. 1987.甘 肃 丫 蕊 花 gan su ya rui huaRhizome 1–2.5 × 0.4–1 cm. Leaves 8 or 9, linear to linearoblanceolate,basally gradually narrowed to a petiole; petiole 2–7 cm × 3–4 mm; leaf blade 3–14 cm × 3–9 mm, apex acuminate.Scape 6–14 cm, usually slightly shorter than basalleaves, densely covered with many imbricate, scaly leaves excepton inflorescence. Raceme 5–13-flowered. Pedicel 3–6 mm,shorter than tepals. Tepals white, tinged purple basally, spatulate-oblanceolate,7–10 × 2–3 mm. Stamens 7–11 mm, slightlyextending beyond tepals at anthesis. Ovary 3-lobed apically.Style 0.9–1.3 cm, longer than stamens; stigma capitate, scarcely3-lobed. Capsule deltoid-obovoid, ca. 1 cm across, much shorterthan persistent tepals. Seeds 4–5 mm. Fl. Mar–Apr, fr. May–Jun.Hillsides; 2000–2100 m. S Gansu.3. Ypsilandra cavaleriei H. Léveillé & Vaniot, Mem. Pontif.Accad. Romana Nuovi Lincei 23: 375. 1905.小 果 丫 蕊 花 xiao guo ya rui huaYpsilandra parviflora F. T. Wang & Tang.Leaves usually more than 10, oblanceolate, basally graduallynarrowed to a petiole; petiole 5–12 cm × 2–4 mm; leafblade 3–13 × 2–4.6 cm, apex acuminate. Scape usually slightlylonger than basal leaves. Raceme 6–10-flowered. Pedicel 4–6mm, nearly as long as tepals. Tepals white or pink, spatulateoblanceolate,4–5 × 1.5–2 mm, 3–5-veined. Stamens 5–6 mm,slightly extending beyond tepals at anthesis. Ovary slightly 3-lobed apically; lobes 1/5–1/4 as long as ovary. Style 6–7 mm,slightly longer than stamens; stigma capitate, slightly or scarcely3-lobed. Capsule broadly ovoid, ca. 2/3 as long as persistenttepals. Seeds ca. 4 mm. Fl. Mar–Apr, fr. Apr–May.Shady places on slopes, hillsides along streams; 1000–1400 m.NW Guangdong, Guangxi, Guizhou, S Hunan.4. Ypsilandra yunnanensis W. W. Smith & Jeffrey, Notes Roy.Bot. Gard. Edinburgh 9: 143. 1916.云 南 丫 蕊 花 yun nan ya rui huaHelonias yunnanensis (W. W. Smith & Jeffrey) N. Tanaka;Ypsilandra yunnanensis var. himalaica H. Hara; Y. yunnanensisvar. micrantha Handel-Mazzetti.Rhizome rather short. Leaves 6 or 7, basally gradually narrowedto a petiole; petiole 0.7–4 cm × 3–5 mm; leaf blade spatulate,2–9 × 1–3.5 cm, apex subacute. Scape 3–30(–40) cm,much longer than basal leaves. Raceme rather narrow, densely5–17-flowered. Pedicel 2–3 mm. Tepals spatulate or oblanceolateto elliptic, 4–5 × 1–2 mm. Stamens 3–4 mm, not extendingbeyond tepals at anthesis or slightly so in fruit. Ovary slightly3-lobed apically. Style 1–1.5 mm, not elongate in fruit; stigmadeeply 3-lobed; lobes ca. 0.8 mm, recurved. Capsule broadlyobovoid, slightly longer than persistent tepals. Seeds ca. 5 mm.Fl. Jun–Jul, fr. Aug–Oct.Rhododendron forests, thicket margins, grassy slopes; 2700–4000(–4300) m. NE Xizang, NW Yunnan [Bhutan, Myanmar, Nepal].5. Ypsilandra alpina F. T. Wang & Tang, Bull. Fan Mem. Inst.Biol. 7: 81. 1936.高 山 丫 蕊 花 gao shan ya rui huaHelonias alpina (F. T. Wang & Tang) N. Tanaka.Leaves several, basally gradually narrowed to a petiole;petiole 3–4 cm; leaf blade lanceolate, 7–8 × 1–1.4 cm, apexacute. Scape 22–26 cm, much longer than basal leaves. Racemerather narrow, 2–6-flowered. Pedicel 3–4 mm. Tepals yellow,linear to linear-lanceolate, 7–12 × 1.5–2 mm. Stamens 4–9 mm,shorter than tepals at anthesis. Ovary slightly 3-lobed apically.Style 2.5–6 mm; stigma shortly to moderately 3-lobed; lobes0.3–0.5 mm. Capsule shorter than tepals. Fl. Jul–Oct.Thicket margins, grasslands; 2000–4300 m. SE Xizang, NWYunnan [Myanmar].7. HELONIOPSIS A. Gray, Mem. Amer. Acad. Arts, ser. 2, 6: 416. 1858, nom. cons.胡 麻 花 属 hu ma hua shuChen Xinqi ( 陈 心 启 Chen Sing-chi); Minoru N. TamuraHexonix Rafinesque, nom. rej.; Kozola Rafinesque, nom. rej.; Sugerokia Miquel.Herbs perennial, with a short, thickened rhizome, glabrous. Leaves basal, rosulate, narrowly oblong to oblanceolate or obovate,proximally gradually narrowed to a petiole, evergreen, margin smooth, sometimes minutely undulate. Scape arising from center ofleaf rosette, erect, simple, hollow, with 2–8 scale leaves. Inflorescence a terminal umbel or umbel-like raceme, 1–10-flowered; bractusually absent. Flowers bisexual, usually slightly nodding at anthesis, ascending in fruit, spreading funnelform. Tepals 6, free, spatulateor linear-oblanceolate to oblong, adaxially often with a deep pocket of a nectary gland at base, persistent. Stamens 6, oftenadnate basally to tepals, sometimes free, always free from ovary, usually exserted, rarely included; anthers lanceolate, dorsifixed,extrorse to latrorse, sometimes with locules confluent at apex. Ovary superior, 3-loculed; ovules 60–180 per locule. Style 1, slender,rather long; stigma capitate. Fruit a capsule, 3-lobed, loculicidal. Seeds small, linear, both ends caudate.Five species: <strong>China</strong>, Japan, Korea; one species (endemic) in <strong>China</strong>.Tanaka (J. Jap. Bot. 73: 102–115. 1998) reduced Heloniopsis and Ypsilandra to synonymy under Helonias. However, a recent molecularphylogenetic study by Fuse and Tamura (Plant Biol. 2: 1–13. 2000) confirmed that Helonias, Heloniopsis, and Ypsilandra are not mixed with eachother, and they each deserve independent generic status.1. Heloniopsis umbellataBaker, J. Bot. 12: 278. 1874. 胡 麻 花 hu ma hua
- Page 3: 34b. Stem very short; leaves basal
- Page 6: simple, erect, usually with a few s
- Page 9 and 10: ciflorus and S. spicatus Klotzsch);
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- Page 16 and 17: Helonias umbellata (Baker) N. Tanak
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- Page 20 and 21: 小 重 楼 xiao chong louPlants 7-
- Page 22 and 23: 14b. Paris thibetica var. apetala H
- Page 24 and 25: slightly convex connective apically
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- Page 30 and 31: 8. Smilax davidiana A. de Candolle
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- Page 36 and 37: slightly shorter than petiole; umbe
- Page 38 and 39: 马 甲 菝 葜 ma jia ba qiaVines
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- Page 42 and 43: 70. Smilax bauhinioides Kunth, Enum
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2b. Flowers not campanulate, tepals
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etween apical whorl and basal bract
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Lilium nanum Klotzsch var. brevisty
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2.5-3 cm × 5-10 mm. Stem 20-30 cm,
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5.5-6 cm × 8-10 mm, margin revolut
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woolly hairs ......................
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3-5-veined, margin papillose. Flowe
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3a. Inner tepals neither spotted no
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3a. Stems subglabrous.4a. Flowers p
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Tepals obliquely outward spreading,
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12a. Tepals white to cream, spotted
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latipetalum Collett & Hemsley; D. p
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only slightly exserted from periant
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laysia, Myanmar, Nepal, Philippines
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3b. Perianth tube longer, to 4 cm,r
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About 660 species: N hemisphere, ma
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44a. Scape 10-15(-20) cm; perianth
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85b. Perianth segments 3-6 mm; fila
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125b. Inner filaments entire at bas
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Leaf blade without white main veins
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irregularly 2-lobed. Filaments subu
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Scrub, moist places; 4500-5000 m. S
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Forests, slopes, pastures; 1600-330
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(40-)50-80 cm, terete, covered with
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2/3-5/6 as long as perianth segment
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Filaments shorter than perianth seg
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umbel, covered with leaf sheaths on
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shores; 1900-3700 m. Xinjiang [Afgh
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昆 仑 韭 kun lun jiuBulbs cluste
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Bulb solitary or clustered, applana
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北 葱 bei congBulbs usually clust
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112. Allium songpanicum J. M. Xu in
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Style exserted. Fl. and fr. May-Jul
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Forests on shady slopes, sands, san
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stigma small. Fruit a capsule, glob
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90: 27. 1931.紫 萼 zi eBryocles v
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Tepals white, linear, membranous; o
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19a. Male flowers 2-4 mm; branches
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perianth yellowish green, campanula
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Asparagus soongoricus Iljin.Herbs d
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北 天 门 冬 bei tian men dongAs
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fr. Sep-Nov.Forests, bamboo forests
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Tovaria oleracea Baker, J. Linn. So
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nate at base, outer ones broadly ov
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2.5 mm. Style ca. 2 mm; stigma capi
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38b. Peduncle usually 1-1.5 cm; inf
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5-12(-17)-flowered; peduncle 3-5 cm
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Himal. Mts. 1: 380. 1839.对 叶
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mm. Perianth pale yellow, cylindric
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nan [Laos, Thailand, Vietnam].2. Di
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Convallaria keiskei Miquel; C. keis
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margin, free part very short. Ovary
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sometimes greenish, ovate to lanceo
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Liang Songyun ( 梁 松 筠 Liang S
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lobes with a white hollow on adaxia
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ar-lanceolate, 30-40 × 3-5 cm, lea
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26. Aspidistra cruciformis Y. Wan &
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Fl. Apr. 2n = 36*.SW Guangxi (Longz
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2.5 mm. Flowers solitary, sometimes
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17b. Plants less than 40 cm tall; t
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asal one 6-8 mm. Flowers solitary;
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2n = 36*, 72*.Dense forests, mossy
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Plants stoloniferous; stolons with
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ulate distally. Tepals purple, or w
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Stem 2-3 cm. Leaves 2 or 3; petiole