New griffenfly, Bohemiatupus elegans from the Late Carboniferous ...

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J. Prokop & A. NelNew insect material was recently discovered atOvčín near Radnice during extensive excavations bypalaeobotanists in tuff layers so called „whetstonehorizon“ from the roof of the Lower Radnice Seam(Lower Bolsovian) with well known buried plantspreserved in situ (see Opluštil et al. 2007, 2009;Libertín et al. 2009). Insect fauna is very rare with twoother undescribed specimens of palaeodictyopteridlarvae and orthopteroid insect (Prokop unpubl.).Material and methodsThe fossil specimen was observed under stereomicroscopes ZeissCytoplast in dry state and under ethyl alcohol. Due to largedimensions of the specimen the venation pattern line drawingwas redrawn from color photograph and revised by parallelobservation under stereomicroscope. Drawing was finallyreadjusted to the photograph scale using of graphic software(Adobe Photoshop). Photographs were made in dry state andunder ethyl alcohol by digital camera Nikon D80 with macrolens Nikon AF-S VR Micro-Nikkor 105 mm in the highestcontrast as possible by single sided cross-light pre-exposure.Type material was collected in 2006 during extensive excavationsby group of palaeobotanists so called “Czech mafia” in Ovčinnear Radnice and housed in the collection of West BohemianMuseum in Plzeň, Czech Republic.We use the wing venation nomenclature of Nel et al. (1993),and Bechly (1996). Systematic and divisions follows the conceptof higher phylogenetic classification of Odonatoptera proposedby Bechly (1996, 2007).The venational symbols used here specified as follows: symbolsin capitals denote the longitudinal veins (CA / CP – costalanterior / posterior, ScA / ScP – subcostal anterior / posterior,RA / RP – radial anterior / posterior, IR – intercalary radialposterior, MA / MP – medial anterior / posterior, CuA / CuP– cubital anterior / posterior, AA – anal anterior).Figure 1Bohemiatupus elegans n. gen., n. sp. (Meganeuridae), holotype M00485 (negative imprint) – line drawing of fore- and hindwing venation, isolated apical partdrawn as preserved.184
New Late Carboniferous griffenfly in western BohemiaResultsSuperorder Odonatoptera Martynov 1932Order Meganisoptera Martynov 1932Family Meganeuridae Handlirsch 1906Subfamily Tupinae Handlirsch 1919Genus Bohemiatupus n. gen.Type species. Bohemiatupus elegans n. sp.Etymology. Composite name after Bohemia (historical regionin central Europe, occupying the western two-thirds of thetraditional Czech Lands, currently the Czech Republic) andTupus (genus name), masculine in gender.Diagnosis. ScA short; ‘subnodus’ present between RA and RPnear base of RP2; parts of CuA and CuP between MP and AAseparated and oblique, with only one cells between them; AA1with posterior branches; CuP very long and straight, with morethan three concave branches, but area covered by posteriorbranches of CuP distinctly narrower than those of CuA andAA1; hind wing distinctly broader and with anal and cubitoanalareas distinctly broader than in forewing.Bohemiatupus elegans n. sp.(Figs. 1–4)Holotype. Specimen M00485, well preserved basal and medialpart of fore- and hindwing, apical part of fore wing distortedand medioapical part of hindwing not preserved, negativeimprint, collection of West Bohemian Museum in Plzeň, CzechRepublic.Figures 2–4Bohemiatupus elegans n. gen., n. sp. (Meganeuridae), holotype M00485 (negative imprint). 2, photograph of wing venation, ventral view; 3, detail photographof basal part of forewing; 4, detail photograph of forewing isolated apical part.185
Page 1: Ann. soc. entomol. Fr. (n.s.), 2010
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New griffenfly, Bohemiatupus elegans from the Late Carboniferous ...

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