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Congressionally classified and designated areas - areas that require congressionalenactment for their establishment, such as national wildernessareas, national wild and scenic rivers, and national recreation areas. RGConifer - a tree belonging to the order Gymnospermae, comprising a widerange of trees that are mostly evergreens. Conifers bear cones (henceconiferous) and needle-shaped or scale-like leaves. SAFConnectivity - a measure of the extent to which intervening habitat trulyconnects DCAs for juvenile spotted owls dispersing among them.Conservation - the process or means of achieving recovery.Conspecifics - belonging to or pertaining to the same species.Consultation - a formal interaction between the U.S. Fish and Wildlife Serviceand another federal agency when it is determined that the agency's actionmay affect a species that has been listed as threatened or endangered orits critical habitat.Contiguous habitat - habitat suitable to support the life needs of owls that isdistributed continuously or nearly continuously across the landscape.Corridor - a defined tract of land, usually linear, through which a species musttravel to reach habitat suitable for reproduction and other life-sustainingneeds.Cost efficiency - the usefulness of specified inputs (costs) to produce specifiedoutputs (benefits). In measuring cost efficiency, some outputs, includingenvironmental, economic, or social impacts, are not assigned monetaryvalues, but are achieved at specified levels in the least costly manner.Cost efficiency usually is measured using present net value, although useof benefit-cost ratios and rates-of-return may be appropriate. RGCritical habitat - specific areas within the geographical area occupied by aspecies on which are found those physical or biological features essentialto conservation of the species.Crown - the upper part of a tree or other woody plant which carries the mainsystem of branches and the foliage. SAFCrown closure - see canopy closure.CWD (coarse woody debris) - portion of a tree that has fallen or been cut andleft in the woods. Usually refers to pieces at least 20 inches in diameter.DBH - diameter at breast height. The diameter of a tree measured 4 feet 6inches from the ground. RGDCA - designated conservation area.Defoliators - insects that feed on foliage and thus act to remove some or all ofthe foliage from a tree, shrub, or herb.Demographic model - a model that predicts the future state of an animalpopulation based on its birth and death rates.Demography - the quantitative analysis of population structure and trends:population dynamics. EFG648
Density, biological population - the number or size of a population in relationto some unit of space. It is usually expressed as the number of individualsor the population biomass per unit area or volume. RGDepauperate - poorly developed. In biology, it usually refers to an area thathas relatively few plant and animal species.Designated conservation area (DCA) - a contiguous area of habitat to bemanaged and conserved for spotted owls. This general description can beapplied to two categories:DCA 1 - category of DCA intended to support at least 20 pairs ofspotted owls.DCA 2 -category of DCA intended to support one to 19 pairs ofspotted owls.Dispersal - the movement, usually one way and on any time scale, of plants oranimals from their point of origin to another location where they subsequentlyproduce offspring.Dispersal capability - ability of members of a species to move from their areaof birth to another suitable location and subsequently breed.Dispersal habitat - habitat that supports the life needs of an individual animalduring dispersal. Generally satisfies needs for foraging, roosting, andprotection from predators.Distribution (of a species) - the spatial arrangement of a species within itsrange.Disturbance - a significant change in structure and/or composition caused bynatural events such as fire and wind or human-caused events such ascutting.Diversity - see biological diversity.Down log - portion of a tree that has fallen or been cut and left in the woods.Early seral stage forests - Stage in forest development that includes seedling,sapling, and pole-sized trees.East side forests - the 12 national forests in Washington, Oregon, and Californiathat lie partly or wholly east of the Cascade Mountain Range crest:Colville, Deschutes, Fremont, Klamath, Malheur, Ochoco, Okanogan,Shasta-Trinity, Umatilla, Wallowa-Whitman, Wenatchee, and WinemaNational Forests. RGEcosystem - an interacting system of organisms considered together with theirenvironment: for example, marsh, watershed, and lake ecosystems. RGEdge - where plant communities meet or where successional stages or vegetativeconditions within plant communities come together. RG See also edgecontrast and horizontal diversity.Edge contrast - a qualitative measure of the difference in structure of twoadjacent vegetated areas; for example, 'low," "medium," or 'high" edgecontrast. RGElectrophoresis - a technique used to distinguish among allozymes by comparingtheir relative mobility in an electrical field.649
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I 1.2:Sp 6/draftRecovery Plan for t
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List of FiguresTable 4.1 Abbreviate
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Recovery ObjectiveSecretary of the
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Management Rules for Designated Con
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Monitoring and Research ProgramThe
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Chapter IIntroduction1
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"Threatened species" means a specie
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B. The Interagency Scientific Commi
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lished works and many government do
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II.A. Natural History of theNorther
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I- \IzXelT - - -.11 I..., ... I.."
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spring. Territories probably are sm
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Northern goshawks (Accipiter gentil
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also use a variety of nest sites in
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Home Range SizeHome range is define
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Table 2.2. Median amounts of old-gr
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3. Life Historycannot reject the hy
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Dispersal1984, Thomas et al. 1990,
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4. ConclusionOur knowledge of the n
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Ca n a d aEastern Washington Cascad
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Table 2.3. Estimated spotted owl ha
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Table 2.4. Significant threats to t
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Table 2.5. Results of surveys for s
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isolated. In these provinces, small
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centers for 30 to 35 spotted owl te
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Southwest Washington occupies a key
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Distribution of Habitat and Populat
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Province Isolation. The eastern Was
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were in suitable habitat condition.
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Estimated acres of forest landbase:
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owned and contain little suitable h
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Vulnerability to Natural Disturbanc
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Douglas-fir/hardwood types, the lat
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miles from the contiguous populatio
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In comparison, the southern part of
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Sacramento River Canyon now provide
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On January 11, 1980, there was an a
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vation Strategy for the Northern Sp
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transfer of one area to the Grand R
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The BLM Spokane (Washington) Distri
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3. National Park ServiceThe followi
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The results of a consultation are s
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- Since landowners already face cut
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4. Cumulative Effects AssessmentThe
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Assessment, Planning, and Monitorin
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Spotted owl nesting sites and activ
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Coastal Oregon Productivity Enhance
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including wildlife. In other cases,
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Currently, the Yakima Indian Nation
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where timber harvest is limited to
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III.A. Recovery Objective and Delis
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4. The population is unlikely to ne
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Needs of other species should be co
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years. The strategy of managing for
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't0
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Finally, the plan recommends mainta
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A total of 1,181 pairs of owls has
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DKnown owlsin the OregonprovincesKn
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mologists, and representatives of o
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this prey species is an appropriate
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After 100 years residual snags will
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2. Fuelwood Gathering. If allowed,
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The application of prescriptions A,
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owls. (Refer to tables in section I
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PRESCRIPTION D -RETAIN OWLS IN MANA
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126
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DCA Management Plans.The recovery p
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Endangered Species Act) or through
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6. Costs to landowners should be re
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The real or perceived disincentives
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authority of the public body to ent
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* Use the recovery plan's recommend
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140
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Supplemental pair areas - Habitat d
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Numbers of owls currently are estim
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Biological goals and implementation
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ning. Also, several unsurveyed area
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Owl pairsKnown owlsin the province3
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- Initiate long-range planning effo
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Table 3.8. Summary comments on the
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from nonfederal lands are needed to
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Eastern Washington Cascades Provinc
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(Owl pairs140 - _ _ _ _ _ _ _ _ _ _
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habitat in DCA WD- 16. Approximatel
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Table 3.12. Summary comments on the
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Ttllwrnook/Astoria area. Provide su
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connectivity between key DCAs. This
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ITable 3.14. Summary comments on th
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crest of the Cascade Mountains. The
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ITable 3.16. Summary comments on th
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sites on federal lands (figure 3.21
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Owl pairsL I 350- _ _ . ...........
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DCAs, often combined with adjacent
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Owl pairsLiKnown owlsin the provinc
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nesting and roosting habitat until
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Option 3: Management of clusters: g
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Table 3.21. Summary of acreage and
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Owl pairs -Known owlsin the provinc
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and to within 0.5 miles of the acti
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Spotted owls have been found at 86
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ITable 3.24. Summary comments on th
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their distance from other sites. Ar
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* Recommend population and habitat
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away in some or all of the range. D
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4. What are the population dynamics
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If the monitoring and research prog
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lowlands, eastern Oregon Cascades,
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210
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Stepdown Outline1. Management Tasks
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Table 4.1 Abbreviated Cost TableThe
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continued-TaskPriorityResp. PartyCo
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218
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220
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and the number of invertebrate anim
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Table 5.2. Numbers of other species
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Amphibians and Reptiles: Larch Moun
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228
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230
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Barrows, C. W. 1980. Feeding ecolog
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Clark, R. J., D. G. Smith, and L. H
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Franklin, A. B., J. A. Blakesley, a
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Harestad, A. S., and F. L. Bunnell.
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Lundquist, R. W. and J. M. Mariani.
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O'Halloran, K. 1989. Spotted owl in
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Solis, D. M. 1980. Habitat use by n
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Young, K. D., A. B. Franklin, and J
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248
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250
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Table A.1. General approaches for e
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complete count of the territorial b
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Table A.3. Powera for various desig
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Table A.4. Illustration of a Markov
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Obviously, these statements hold on
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Table A.6. Reliability of trend est
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Demographic AnalysisAs noted earlie
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the estimate (using the simple equa
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however, we know little about the b
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Table A.8. Summary of information n
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predict the power achieved by diffe
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Other studiestached. If dispersing
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impractical at present. We describe
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Conclusionsthe year effect can be i
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280
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282
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284
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owls. Consequently, the issue of wh
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Use of the informationAssessments a
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Recent LiteratureWe might begin by
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Kerms (1989) measured habitat struc
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Hamer (pers. comm.) described 11 ne
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Table B.3. Percent hardwoods in the
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Table B.5. Comparison of habitats u
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Table B.8. Tree density (number of
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and numbers of trees, and percent o
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Under this hypothesis, owl fitness
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suitable habitat for that region (m
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ResultsDiscussionspotted owl habita
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__Discussiondata from 41 sites in t
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Recommendations for Future Research
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314
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Hamer, T. 1988. Home range size of
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318
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320
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Parameter Estimatesfor Individual S
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provided in Table C. 5. Study of th
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Table C.6. Summary of statistics re
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3. ConclusionsThis source of bias i
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330
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Gray Wolf .........................
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formed during the initial meeting t
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provide habitat for the numerous sp
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The ApproachA committee of the Reco
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The List of Threatened and Endanger
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Mammalsoldest trees in a stand. Eig
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ABBFigure D. 1. Distribution of: a)
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KTable D.3. A list of threatened an
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Fishestheir dispersal capabilities
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OlympicNorthcoastalRockyMountainSou
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continued-StatusabState Considered
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continued-SpeciesColumbia sidebandM
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endemic to the Pacific Northwest).
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------continued-SpeciesRhyacophila
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continued-SpeciesFed3tatusa bStateW
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StatusasbOld ForestState Associatio
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Summaryspores (Maser et al. 1978b).
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Biology of Owl Prey, Older Forest A
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Woodrats.-The dusky-footed woodrat
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and are less restrictive under stat
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Stock concept.-The term "stock" was
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Table D.10. List of fish stocks of
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continued-Steelhead-owipu Sea-runTr
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continued-BasinHoodWillametteRogueS
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Table D.13. Washington streams (or
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inland distribution currently decre
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nesting territories in the three st
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However, little is known about the
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Fisher (Martes penncmti)Distributio
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at one time supported marten popula
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each population of these animals is
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species also is found under the bar
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few meters from water after heavy r
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In considering the needs of other s
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Barrows, C. W. 1981. Roost selectio
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Clark, T. W., E. Anderson, C. Dougl
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Hamer, T. E. 1991. Habitat relation
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Mannan, R. W., E. C. Meslow, and H.
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Oakley, A. L., Collins, J. A., Ever
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Solis, D. M. 1983. Summer habitat e
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Washington Natural Heritage Program
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414
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f. Wilderness.g. Livestock grazing.
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I. Monitoring.1. Northern spotted o
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420
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here will reduce some of that habit
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424
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426
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Selection of Forest TypesAt the tim
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eceive as little as 25 inches. Disc
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A. FireThe combination and interact
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Table F.4. Important forest insects
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continued-Common Name Causal Agent
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D. DiseasesForest diseases in the P
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continued-Common Name Causal Agent
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continued-Common Name Causal Agent
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III. Forest Protection in theWest C
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120 100200 500I~~~~.K446Figure F.4.
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InsectsDiseasesInsects in this subr
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__hardwoods are immune. Shade toler
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pests (Schowalter 1988). Black stai
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tion and initial attack. If manipul
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log and snag density was likely low
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B. Management Effects on StandsMana
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units (underburning or lop/scatter)
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In the higher elevation White Fir a
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growth often increases (Reinhardt a
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50rcc4*0-cC)2M CmZ13010-yearI ,...*
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In the East Cascades subregion fore
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The introduced disease, white pine
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VI. Conclusionsmortality will resul
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Baker, F.A. 1988. The influence of
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Habeck, J.R. 1990. Old-growth ponde
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Perry, D.A., and J.G. Borchers. 199
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Weatherspoon, C.P., and C.N. Skinne
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482
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484
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area, structure, ages, location; ne
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Stand AFigure G.2. A--unthmnned Dou
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figure 0.3. Effects of thinning Dou
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II. Review of Natural Vegetation an
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western hemlock, and sometimes othe
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year intervals (Appendix F). With t
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Live crown ratiobecent)6050IStand A
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Basal area growth continued to incr
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technology probably will allow some
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Stand A. No activities (west side 3
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Stand A. No activities (west side o
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acre, 4 to 12 inches in diameter),
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allow redwood sprout clumps maximum
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KAge 15 years - 511 trees per acre,
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6Stems per acre100 _ Stand A. Natur
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Basal Area(square feetper acre)50 1
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* Opening the canopy in small patch
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Drew, T.J. and J.W. Flewelling. 197
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Miller, R.E.; D.L. Reukema, and R.L
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Tappeiner, J.C., P.M. McDonald, and
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526
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528
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6. Timber supply curve and economic
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scientifically credible biological
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E. Summary of Preliminary Estimates
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II. General Economic Principles for
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practices may be able to show how s
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Economic patterns show the efficien
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The sustainable yield concept that
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A. Previously Estimated Economic Co
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County receipts were estimated to d
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B. Potential Social Costs of Owl Co
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The contrast between the behavior d
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0U1 n1nDTable H.2. Economic depende
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cḻlTable H.4. Economic characteri
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An analysis completed by the State
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V. Preliminary Evaluation of the Ec
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25-° 20/0M 15-a)CLU)0 10-0CL 5 -0)
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25 -- 20-0co/o 15--Da)a0a)10m~ C) 5
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forestlands. The costs of productio
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In Figure H.8, the equilibrium betw
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foregone and the way they will save
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The income created by the purchase
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Two concepts are useful for underst
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y an estimate of the price that wou
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Table H.10. Comparison of timber em
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of the second year. The same assump
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are generally much lower than for c
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582
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studies at University of Washington
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586
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Most significant late successionall
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to evaluate options. Information in
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Q0Table 1.1. Estimated acres and ow
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soTable 1.3. Estimated acres and ow
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01CDarTable 1.5. Size class distrib
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Table 1.8. Frequency distribution o
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Literature CitedThomas, J.W., E.D.
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602
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- The overlap of home ranges among
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Table J.2. Acreage and owl numbers
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continuedDCA Land AcreageIdent. Sta
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Table J.4. Acreage and owl numbers
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Table J.5. Acreage and owl numbers
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Table J.6. Acreage and owl numbers
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Table J.7. Acreage and owl numbers
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Table J.9. Acrea ?e and owl numbers
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Table J.10. Acreage and owl numbers
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continued-DCA LandIdent. StatusAcre
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-DCAIdent.LandStatusAcreageNRF Tota
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626
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Experience Includes: Assistant Dire
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Education: B.S. in forestry from Or
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Team SupportCharles Bruce, Oregon D
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continued-1 5-24-91 Other SpeciesSu
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continued-9-16 thru9-20-919-19-91Sp
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Perry, D. 1991. The ecology of coar
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2mandate the achievement of recover
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642
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644
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Aspect - the direction a slope face
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Empirical - derived from direct obs
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Hummocky - a landscape characterize
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Mixed conifer - as used in this doc
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Potential habitat - a stand of tree
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Sensitive species - those species t
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Take -Uunder the Endangered Species
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wild, scenic, or recreational by an
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