Recovery Plan for the Northern Spotted Owl - DRAFT
Recovery Plan for the Northern Spotted Owl - DRAFT Recovery Plan for the Northern Spotted Owl - DRAFT
Basal area growth continued to increase for 70 years, indicating that trees werevigorous and continued to respond to the growing space provided by the lowstocking. An understory of bigleaf maple, grand fir, and shrubs (e.g., hazel andoceanspray) had developed in these stands.Windthrow is a natural part of stand development which will occur in thinnedand unthinned stands. It is not necessarily a concern after thinning, even inolder stands. Graham et al. (1985) recorded windthrow on 400 acres ofthinned 100-year-old Sitka spruce-western hemlock stands near the coast atCascade Head, Oregon, from which 0 to 25 percent of the basal area wasremoved. Windthrow was variable and occurred in thinned and unthinnedsites. For example, all windthrow in one tract occurred on half the plots, andtracts which had heavy windthrow during some storms had only very lightlosses during others. Overall volume of windthrown trees was nearly twice asgreat in unthinned stands as it was in thinned stands, possibly because treesprone to wind damage were removed in thinning. Total mortality from allcauses, but not including thinned volume, was somewhat less on thinned sites(225 cubic feet per acre) than on unthinned sites (300 cubic feet per acre); thisamounts to about only one year's volume growth. Similarly, Williams and Price(1971), after studying 70- to 150-year-old stands 18 years after thinning,concluded that windthrow was related to factors such as soil depth andtopographic position and wind patterns rather than thinning.Ruth and Harris (1979) discussed the problems of wind and thinning inwestern hemlock-Sitka spruce forests. They concluded that site (topography,soil depth, soil drainage), root disease, type of thinning, and age at thinning (orother disturbance) predispose stands to windthrow. Thinning old, densestands likely will increase risk of damage on windy sites, while thinningyounger stands may decrease the risk at later stages of development. Leavingwind firm edges that do not channel wind and unthinned buffers is importanton exposed sites because they are likely to decrease the risk of blowdownprogressing throughout a stand. Thinning also lowers height to diameterratios of trees by increasing stem diameter growth relative to height growth andby lowering the center of gravity of the crown. Based on European experience,height to diameter ratios of 100 or more in Douglas-fir indicate susceptibility todamage (Worthington and Staebler 1961). Thinning would decrease this ratiobecause tree diameter growth usually is increased by thinning more thanheight growth is increased.B. Understory Vegetation and Advanced Regeneration500Regulating stand density affects understory vegetation. As the overstorybecomes less dense from thinning or natural disturbance, an understory ofshrubs, hardwoods, conifers, and other species develops. Conifer and hardwoodseedlings and saplings will grow to provide the multilayered stands usedby owls. Alaback and Herman (1988) reported the development of a twolayeredconifer stand 17 years following thinning in western hemlock andspruce stands. However, development of too dense an understory of hemlockmay inhibit development of shrub or forb layers. After disturbance to theoverstory (release), conifer seedlings and saplings increase their growth, and itappears that in open conditions the rate of growth following release is relatedto the size of their crowns and rates of prerelease height growth (Helms andSandiford 1985, Gordon 1973, Stein 1981, 1986). Careful removal of overstorytrees is needed to ensure release of advanced regeneration (Mann and Tesch1985, Tesch et al. 1990a); however, Douglas-fir seedlings can recover fromdamage by overstory removal (Tesch et al. 1990b). Growth of western hemlockand western red cedar present in the understory of a recently thinned 110-year-old Douglas-fir stand was increased by fertilization or thinning(Harrington and Wierman 1990).
C. FertilizationThinning or other disturbance also aids the development of shrubs (Long andTurner 1975) (see Figure G.3). Vine maple clones spread by layering as theyare pinned to the ground by falling trees (O'Dea 1991). Shrubs like salmonberryand salal spread from increased rhizome extension and seedling establishmentin thinned stands flappeiner et al. 1991, Huffman 1991). Too densea layer of shrubs or forbs will prevent the establishment of conifers and hardwoodsin the understory (Isaac 1938). Bigleaf maple seedlings become establishedas the overstory is thinned to about 70 to 80 percent cover or less, butbefore a layer of shrubs or forbs is formed (Fried et al. 1988). Where layers ofshrubs occur in the understory, some treatment to reduce their density may beneeded to establish trees to provide the stand structure used by owls.Fertilization has been shown to increase the rate of stand development instands 15 to 110 years old. For example, 90-year-old (Miller and Harrington1979) and 110-year-old (Harrington and Miller 1979) Douglas-fir standsincreased basal area and diameter growth rates following application of nitrogenfertilizer as urea or ammonium nitrate. Increased growth occurred inthinned and unthinned stands and on a range of sites and stand ages lessthan 90 years (Miller et al. 1979, Miller and Tarrant 1983). Fertilization alsoincreased diameter and height growth in thinned and unthinned 15-year-oldwestern red cedar stands (Harrington and Wierman 1990).D. Historical Stand Management PracticesThe primary silvicultural system for Douglas-fir and western hemlock forests inthe coastal forests and western Cascades of Oregon and Washington has beeneven-age management. This is an efficient way to manage these forests fortimber production primarily because a) the major conifers can be regeneratedin open conditions following disturbance, b) steep topography on many sitesprecludes tractor logging or makes it prohibitively expensive and likely to causeserious soil erosion, and c) cost of setting up cable logging equipment and roadbuilding in steep terrain makes it most efficient and least expensive to clearcut.Under the even-age system, stands most often are regenerated using theclear-cutting method, followed by site preparation, planting nursery-grownseedlings, and some level of control of herbs, shrubs, or hardwoods. Theshelterwood method also has been used on some sites where wind is not aserious problem (Williamson 1973).Early in the development of silvicultural practices in this region, Kirkland andBrandstrom (1936) recommended partial cutting and group selection as a wayto manage Douglas-fir. Isaac (1956) reviewed the early trials of this type of"selection management" in old-growth forests and recommended that Douglasfirforests be managed under even-age systems. His recommendations werebased on observations that a) there was little natural regeneration occurringafter selection cutting, b) understory trees were damaged by logging, c) therewas blowdown (few trees, but lots of volume), and d) the old trees generally didnot increase their growth rates after cutting. Isaac (1956) recognized severalcases where selection cutting (uneven-age management) appeared to be reasonable:a) dry sites with more open stands, b) stands with a high proportionof young trees, and c) salvage of mortality. It is important to note that hisrecommendations were based mainly on experience in old stands, not youngstands (less than 100 to 120 years). These recommendations considered onlynatural regeneration without well-planned site preparation prior to planting ornatural seeding, followed by vegetation control. Also, advances in logging501
- Page 466 and 467: __hardwoods are immune. Shade toler
- Page 468 and 469: pests (Schowalter 1988). Black stai
- Page 470 and 471: tion and initial attack. If manipul
- Page 472 and 473: log and snag density was likely low
- Page 474 and 475: B. Management Effects on StandsMana
- Page 476 and 477: units (underburning or lop/scatter)
- Page 478 and 479: In the higher elevation White Fir a
- Page 480 and 481: growth often increases (Reinhardt a
- Page 482 and 483: 50rcc4*0-cC)2M CmZ13010-yearI ,...*
- Page 484 and 485: In the East Cascades subregion fore
- Page 486 and 487: The introduced disease, white pine
- Page 488 and 489: VI. Conclusionsmortality will resul
- Page 490 and 491: Baker, F.A. 1988. The influence of
- Page 492 and 493: Habeck, J.R. 1990. Old-growth ponde
- Page 494 and 495: Perry, D.A., and J.G. Borchers. 199
- Page 496 and 497: Weatherspoon, C.P., and C.N. Skinne
- Page 498 and 499: 482
- Page 500 and 501: 484
- Page 502 and 503: area, structure, ages, location; ne
- Page 504 and 505: Stand AFigure G.2. A--unthmnned Dou
- Page 506 and 507: figure 0.3. Effects of thinning Dou
- Page 508 and 509: II. Review of Natural Vegetation an
- Page 510 and 511: western hemlock, and sometimes othe
- Page 512 and 513: year intervals (Appendix F). With t
- Page 514 and 515: Live crown ratiobecent)6050IStand A
- Page 518 and 519: technology probably will allow some
- Page 520 and 521: Stand A. No activities (west side 3
- Page 522 and 523: Stand A. No activities (west side o
- Page 524 and 525: acre, 4 to 12 inches in diameter),
- Page 526 and 527: allow redwood sprout clumps maximum
- Page 528 and 529: KAge 15 years - 511 trees per acre,
- Page 530 and 531: 6Stems per acre100 _ Stand A. Natur
- Page 532 and 533: Basal Area(square feetper acre)50 1
- Page 534 and 535: * Opening the canopy in small patch
- Page 536 and 537: Drew, T.J. and J.W. Flewelling. 197
- Page 538 and 539: Miller, R.E.; D.L. Reukema, and R.L
- Page 540 and 541: Tappeiner, J.C., P.M. McDonald, and
- Page 542 and 543: 526
- Page 544 and 545: 528
- Page 546 and 547: 6. Timber supply curve and economic
- Page 548 and 549: scientifically credible biological
- Page 550 and 551: E. Summary of Preliminary Estimates
- Page 552 and 553: II. General Economic Principles for
- Page 554 and 555: practices may be able to show how s
- Page 556 and 557: Economic patterns show the efficien
- Page 558 and 559: The sustainable yield concept that
- Page 560 and 561: A. Previously Estimated Economic Co
- Page 562 and 563: County receipts were estimated to d
- Page 564 and 565: B. Potential Social Costs of Owl Co
C. FertilizationThinning or o<strong>the</strong>r disturbance also aids <strong>the</strong> development of shrubs (Long andTurner 1975) (see Figure G.3). Vine maple clones spread by layering as <strong>the</strong>yare pinned to <strong>the</strong> ground by falling trees (O'Dea 1991). Shrubs like salmonberryand salal spread from increased rhizome extension and seedling establishmentin thinned stands flappeiner et al. 1991, Huffman 1991). Too densea layer of shrubs or <strong>for</strong>bs will prevent <strong>the</strong> establishment of conifers and hardwoodsin <strong>the</strong> understory (Isaac 1938). Bigleaf maple seedlings become establishedas <strong>the</strong> overstory is thinned to about 70 to 80 percent cover or less, butbe<strong>for</strong>e a layer of shrubs or <strong>for</strong>bs is <strong>for</strong>med (Fried et al. 1988). Where layers ofshrubs occur in <strong>the</strong> understory, some treatment to reduce <strong>the</strong>ir density may beneeded to establish trees to provide <strong>the</strong> stand structure used by owls.Fertilization has been shown to increase <strong>the</strong> rate of stand development instands 15 to 110 years old. For example, 90-year-old (Miller and Harrington1979) and 110-year-old (Harrington and Miller 1979) Douglas-fir standsincreased basal area and diameter growth rates following application of nitrogenfertilizer as urea or ammonium nitrate. Increased growth occurred inthinned and unthinned stands and on a range of sites and stand ages lessthan 90 years (Miller et al. 1979, Miller and Tarrant 1983). Fertilization alsoincreased diameter and height growth in thinned and unthinned 15-year-oldwestern red cedar stands (Harrington and Wierman 1990).D. Historical Stand Management PracticesThe primary silvicultural system <strong>for</strong> Douglas-fir and western hemlock <strong>for</strong>ests in<strong>the</strong> coastal <strong>for</strong>ests and western Cascades of Oregon and Washington has beeneven-age management. This is an efficient way to manage <strong>the</strong>se <strong>for</strong>ests <strong>for</strong>timber production primarily because a) <strong>the</strong> major conifers can be regeneratedin open conditions following disturbance, b) steep topography on many sitesprecludes tractor logging or makes it prohibitively expensive and likely to causeserious soil erosion, and c) cost of setting up cable logging equipment and roadbuilding in steep terrain makes it most efficient and least expensive to clearcut.Under <strong>the</strong> even-age system, stands most often are regenerated using <strong>the</strong>clear-cutting method, followed by site preparation, planting nursery-grownseedlings, and some level of control of herbs, shrubs, or hardwoods. Theshelterwood method also has been used on some sites where wind is not aserious problem (Williamson 1973).Early in <strong>the</strong> development of silvicultural practices in this region, Kirkland andBrandstrom (1936) recommended partial cutting and group selection as a wayto manage Douglas-fir. Isaac (1956) reviewed <strong>the</strong> early trials of this type of"selection management" in old-growth <strong>for</strong>ests and recommended that Douglasfir<strong>for</strong>ests be managed under even-age systems. His recommendations werebased on observations that a) <strong>the</strong>re was little natural regeneration occurringafter selection cutting, b) understory trees were damaged by logging, c) <strong>the</strong>rewas blowdown (few trees, but lots of volume), and d) <strong>the</strong> old trees generally didnot increase <strong>the</strong>ir growth rates after cutting. Isaac (1956) recognized severalcases where selection cutting (uneven-age management) appeared to be reasonable:a) dry sites with more open stands, b) stands with a high proportionof young trees, and c) salvage of mortality. It is important to note that hisrecommendations were based mainly on experience in old stands, not youngstands (less than 100 to 120 years). These recommendations considered onlynatural regeneration without well-planned site preparation prior to planting ornatural seeding, followed by vegetation control. Also, advances in logging501