Recovery Plan for the Northern Spotted Owl - DRAFT

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eceive as little as 25 inches. Discussion of disturbance and forest protectionfor this zone is most applicable to the wetter plant associations (those withmore than about 60 inches of annual precipitation). The drier western hemlockplant associations are on an environmental gradient toward the Mixed-Conifer Zone discussed for the Klamath subregion, environmentally if not geographically.The Klamath subregion forests include those of the Mixed-Evergreen Zone, theMixed-Conifer Zone, and the White Fir Zone, the latter of which is a moist andcool transition normally above either of the other two zones in southern Oregonand northern California. Douglas-fir is a common dominant in both foresttypes. In the Mixed-Conifer Zone, ponderosa pine, white or grand fir, sugarpine (Pinus lanbertnaro)i or incense-cedar (Calocedrus decurrens) may becodominants. In the Mixed-Evergreen Zone, madrone (Arbutus menziesll),tanoak (Lithocarpus densiflorus), black oak (Quercus kelloggio, and canyon liveoak (Quercus chrysolepis) are common lower canopy dominants, while in theMixed-Conifer Zone, white fir is commonly the lower canopy dominant.The East Cascades subregion includes the mid-elevation Douglas-fir andGrand Fir Zones sandwiched between higher elevation subalpine forest andlower elevation ponderosa pine forests. In some locations, particularly northernWashington, the Grand Fir Zone is absent. Douglas-fir tends to be acanopy dominant, sometimes with western larch (Larix occidentalis) and/orponderosa pine (Pinus ponderosa). Grand fir in the Grand Fir Zone, and Douglas-firin the Douglas-fir Zone, are the most shade-tolerant species and maybe well represented in the lower canopy layers.fX0). Mied @*.@. Ponderosa----- evergreen.'*...Western *i6.- - |.*.9 .............pine'hemlock a . .... Mixed ,'Douglas-fir: . o . g . . f coniferwoodlnd)Figure F.2. Fire severity regimes of the Pac(ficNorthwest. MqJorforest types are ordinated ontemperature and moisture axes, and thendefined as high, moderate, or low severityregimes. The types enclosed by the stippledpattern are those where the northern spottedowl isfound0I-0)0)a-E0)-0-Pacific'silver fir wSubalpineforestsRedfirMoisture stress indexI- -........ "I'l l, I - ......... 1.1.11, 11-1- ..- I ......................430
II. Major Natural Disturbance Factorsof the RegionTraditional theories of disturbance in ecosystems have held that disturbancemust be a major event and must originate outside the ecosystem (i.e., be exogenous)(White 1979). We now embrace a much broader concept of disturbance,recognizing a disturbance gradient from minor to major and the endogenouscomponent of many disturbances (due either to biotic agents orecosystem states that encourage disturbance to occur). As we accept thisbroader concept, we thereby create a fuzzier image of disturbance.Disturbance is a difficult word to define. A simple definition is to "interrupt" or"break up a quiet or settled order." Forest ecosystems, we know, are not "quietor settled orders," but when do the natural, normal rhythms of the systemoscillate to a point where they become "abnormal"? There is no easy answer,but White and Pickett (1985) suggest that disturbance can be defined as ". . .any relatively discrete event in time that disrupts ecosystem, community, orpopulation structure and changes resources, substrate availability, or thephysical environment." It may be characterized by such descriptors as frequency,magnitude, extent, predictability, synergism, and timing (White andPickett 1985). It may include processes such as forest fire, wind, insect, anddisease outbreaks, ice and freeze damage, landslides, floods, and others. Disturbanceneed not be either a disaster or a catastrophe.In Pacific Northwest forests, the primary disturbance factors of concern in DCAprotection are fire, wind, insects, and disease. This report therefore focuses onthese and omits the minor disturbance factors that normally are of limitedextent and major disturbance factors such as volcanoes.I ^-HighFigure F.3. A range offire severity levelsis found within eachfire regime depictedModeratein Figure F.2. The proportion of the-s -_ _ svariousfire severity levels are shown by"' o H ~ s / the lines of the graph. Asfire severityregimes move on the left-to-right gradientfrom low to high, the proportion of lowo / 40 4, severityfires decrease while high severity._fires increase. Moderate severityfireso /.;0 peak in the moderate severity regime.Q ., 1*/ which is the most complexfire regime in. , terms offire effects.Low Moderate HighSeverityType of fire regime- -.......................... - I..431
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I 1.2:Sp 6/draftRecovery Plan for t
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List of FiguresTable 4.1 Abbreviate
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Recovery ObjectiveSecretary of the
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Management Rules for Designated Con
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Monitoring and Research ProgramThe
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Chapter IIntroduction1
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"Threatened species" means a specie
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B. The Interagency Scientific Commi
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lished works and many government do
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II.A. Natural History of theNorther
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I- \IzXelT - - -.11 I..., ... I.."
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spring. Territories probably are sm
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Northern goshawks (Accipiter gentil
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also use a variety of nest sites in
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Home Range SizeHome range is define
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Table 2.2. Median amounts of old-gr
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3. Life Historycannot reject the hy
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Dispersal1984, Thomas et al. 1990,
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4. ConclusionOur knowledge of the n
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Ca n a d aEastern Washington Cascad
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Table 2.3. Estimated spotted owl ha
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Table 2.4. Significant threats to t
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Table 2.5. Results of surveys for s
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isolated. In these provinces, small
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centers for 30 to 35 spotted owl te
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Southwest Washington occupies a key
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Distribution of Habitat and Populat
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Province Isolation. The eastern Was
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were in suitable habitat condition.
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Estimated acres of forest landbase:
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owned and contain little suitable h
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Vulnerability to Natural Disturbanc
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Douglas-fir/hardwood types, the lat
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miles from the contiguous populatio
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In comparison, the southern part of
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Sacramento River Canyon now provide
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On January 11, 1980, there was an a
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vation Strategy for the Northern Sp
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transfer of one area to the Grand R
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The BLM Spokane (Washington) Distri
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3. National Park ServiceThe followi
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The results of a consultation are s
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- Since landowners already face cut
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4. Cumulative Effects AssessmentThe
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Assessment, Planning, and Monitorin
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Spotted owl nesting sites and activ
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Coastal Oregon Productivity Enhance
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including wildlife. In other cases,
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Currently, the Yakima Indian Nation
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where timber harvest is limited to
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III.A. Recovery Objective and Delis
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4. The population is unlikely to ne
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Needs of other species should be co
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years. The strategy of managing for
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't0
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Finally, the plan recommends mainta
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A total of 1,181 pairs of owls has
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DKnown owlsin the OregonprovincesKn
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mologists, and representatives of o
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this prey species is an appropriate
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After 100 years residual snags will
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2. Fuelwood Gathering. If allowed,
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The application of prescriptions A,
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owls. (Refer to tables in section I
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PRESCRIPTION D -RETAIN OWLS IN MANA
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126
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DCA Management Plans.The recovery p
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Endangered Species Act) or through
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6. Costs to landowners should be re
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The real or perceived disincentives
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authority of the public body to ent
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* Use the recovery plan's recommend
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140
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Supplemental pair areas - Habitat d
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Numbers of owls currently are estim
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Biological goals and implementation
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ning. Also, several unsurveyed area
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Owl pairsKnown owlsin the province3
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- Initiate long-range planning effo
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Table 3.8. Summary comments on the
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from nonfederal lands are needed to
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Eastern Washington Cascades Provinc
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(Owl pairs140 - _ _ _ _ _ _ _ _ _ _
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habitat in DCA WD- 16. Approximatel
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Table 3.12. Summary comments on the
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Ttllwrnook/Astoria area. Provide su
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connectivity between key DCAs. This
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ITable 3.14. Summary comments on th
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crest of the Cascade Mountains. The
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sites on federal lands (figure 3.21
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Owl pairsL I 350- _ _ . ...........
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DCAs, often combined with adjacent
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Owl pairsLiKnown owlsin the provinc
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nesting and roosting habitat until
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Option 3: Management of clusters: g
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Table 3.21. Summary of acreage and
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Owl pairs -Known owlsin the provinc
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and to within 0.5 miles of the acti
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Spotted owls have been found at 86
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their distance from other sites. Ar
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* Recommend population and habitat
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away in some or all of the range. D
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4. What are the population dynamics
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If the monitoring and research prog
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lowlands, eastern Oregon Cascades,
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210
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Stepdown Outline1. Management Tasks
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Table 4.1 Abbreviated Cost TableThe
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continued-TaskPriorityResp. PartyCo
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218
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220
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and the number of invertebrate anim
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Table 5.2. Numbers of other species
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Amphibians and Reptiles: Larch Moun
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228
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230
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Barrows, C. W. 1980. Feeding ecolog
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Clark, R. J., D. G. Smith, and L. H
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Franklin, A. B., J. A. Blakesley, a
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Harestad, A. S., and F. L. Bunnell.
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Lundquist, R. W. and J. M. Mariani.
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O'Halloran, K. 1989. Spotted owl in
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Solis, D. M. 1980. Habitat use by n
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Young, K. D., A. B. Franklin, and J
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248
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250
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Table A.1. General approaches for e
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complete count of the territorial b
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Table A.3. Powera for various desig
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Table A.4. Illustration of a Markov
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Obviously, these statements hold on
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Table A.6. Reliability of trend est
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Demographic AnalysisAs noted earlie
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the estimate (using the simple equa
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however, we know little about the b
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Table A.8. Summary of information n
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predict the power achieved by diffe
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Other studiestached. If dispersing
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impractical at present. We describe
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Conclusionsthe year effect can be i
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280
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282
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284
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owls. Consequently, the issue of wh
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Use of the informationAssessments a
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Recent LiteratureWe might begin by
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Kerms (1989) measured habitat struc
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Hamer (pers. comm.) described 11 ne
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Table B.3. Percent hardwoods in the
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Table B.5. Comparison of habitats u
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Table B.8. Tree density (number of
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and numbers of trees, and percent o
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Under this hypothesis, owl fitness
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suitable habitat for that region (m
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ResultsDiscussionspotted owl habita
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__Discussiondata from 41 sites in t
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Recommendations for Future Research
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314
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Hamer, T. 1988. Home range size of
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318
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320
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Parameter Estimatesfor Individual S
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provided in Table C. 5. Study of th
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Table C.6. Summary of statistics re
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3. ConclusionsThis source of bias i
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330
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Gray Wolf .........................
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formed during the initial meeting t
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provide habitat for the numerous sp
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The ApproachA committee of the Reco
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The List of Threatened and Endanger
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Mammalsoldest trees in a stand. Eig
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ABBFigure D. 1. Distribution of: a)
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KTable D.3. A list of threatened an
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Fishestheir dispersal capabilities
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OlympicNorthcoastalRockyMountainSou
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continued-StatusabState Considered
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continued-SpeciesColumbia sidebandM
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endemic to the Pacific Northwest).
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------continued-SpeciesRhyacophila
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continued-SpeciesFed3tatusa bStateW
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StatusasbOld ForestState Associatio
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Summaryspores (Maser et al. 1978b).
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Biology of Owl Prey, Older Forest A
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Woodrats.-The dusky-footed woodrat
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and are less restrictive under stat
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Stock concept.-The term "stock" was
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Table D.10. List of fish stocks of
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continued-Steelhead-owipu Sea-runTr
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continued-BasinHoodWillametteRogueS
Page 396 and 397: Table D.13. Washington streams (or
Page 398 and 399: inland distribution currently decre
Page 400 and 401: nesting territories in the three st
Page 402 and 403: However, little is known about the
Page 404 and 405: Fisher (Martes penncmti)Distributio
Page 406 and 407: at one time supported marten popula
Page 408 and 409: each population of these animals is
Page 410 and 411: species also is found under the bar
Page 412 and 413: few meters from water after heavy r
Page 414 and 415: In considering the needs of other s
Page 416 and 417: Barrows, C. W. 1981. Roost selectio
Page 418 and 419: Clark, T. W., E. Anderson, C. Dougl
Page 420 and 421: Hamer, T. E. 1991. Habitat relation
Page 422 and 423: Mannan, R. W., E. C. Meslow, and H.
Page 424 and 425: Oakley, A. L., Collins, J. A., Ever
Page 426 and 427: Solis, D. M. 1983. Summer habitat e
Page 428 and 429: Washington Natural Heritage Program
Page 430 and 431: 414
Page 432 and 433: f. Wilderness.g. Livestock grazing.
Page 434 and 435: I. Monitoring.1. Northern spotted o
Page 436 and 437: 420
Page 438 and 439: here will reduce some of that habit
Page 440 and 441: 424
Page 442 and 443: 426
Page 444 and 445: Selection of Forest TypesAt the tim
Page 448 and 449: A. FireThe combination and interact
Page 450 and 451: Table F.4. Important forest insects
Page 452 and 453: continued-Common Name Causal Agent
Page 454 and 455: D. DiseasesForest diseases in the P
Page 460 and 461: III. Forest Protection in theWest C
Page 462 and 463: 120 100200 500I~~~~.K446Figure F.4.
Page 464 and 465: InsectsDiseasesInsects in this subr
Page 466 and 467: __hardwoods are immune. Shade toler
Page 468 and 469: pests (Schowalter 1988). Black stai
Page 470 and 471: tion and initial attack. If manipul
Page 472 and 473: log and snag density was likely low
Page 474 and 475: B. Management Effects on StandsMana
Page 476 and 477: units (underburning or lop/scatter)
Page 478 and 479: In the higher elevation White Fir a
Page 480 and 481: growth often increases (Reinhardt a
Page 482 and 483: 50rcc4*0-cC)2M CmZ13010-yearI ,...*
Page 484 and 485: In the East Cascades subregion fore
Page 486 and 487: The introduced disease, white pine
Page 488 and 489: VI. Conclusionsmortality will resul
Page 490 and 491: Baker, F.A. 1988. The influence of
Page 492 and 493: Habeck, J.R. 1990. Old-growth ponde
Page 494 and 495: Perry, D.A., and J.G. Borchers. 199
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Weatherspoon, C.P., and C.N. Skinne
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482
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484
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area, structure, ages, location; ne
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Stand AFigure G.2. A--unthmnned Dou
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figure 0.3. Effects of thinning Dou
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II. Review of Natural Vegetation an
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western hemlock, and sometimes othe
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year intervals (Appendix F). With t
Page 514 and 515:
Live crown ratiobecent)6050IStand A
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Basal area growth continued to incr
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technology probably will allow some
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Stand A. No activities (west side 3
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Stand A. No activities (west side o
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acre, 4 to 12 inches in diameter),
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allow redwood sprout clumps maximum
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KAge 15 years - 511 trees per acre,
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6Stems per acre100 _ Stand A. Natur
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Basal Area(square feetper acre)50 1
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* Opening the canopy in small patch
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Drew, T.J. and J.W. Flewelling. 197
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Miller, R.E.; D.L. Reukema, and R.L
Page 540 and 541:
Tappeiner, J.C., P.M. McDonald, and
Page 542 and 543:
526
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528
Page 546 and 547:
6. Timber supply curve and economic
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scientifically credible biological
Page 550 and 551:
E. Summary of Preliminary Estimates
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II. General Economic Principles for
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practices may be able to show how s
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Economic patterns show the efficien
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The sustainable yield concept that
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A. Previously Estimated Economic Co
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County receipts were estimated to d
Page 564 and 565:
B. Potential Social Costs of Owl Co
Page 566 and 567:
The contrast between the behavior d
Page 568 and 569:
0U1 n1nDTable H.2. Economic depende
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cḻlTable H.4. Economic characteri
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An analysis completed by the State
Page 574 and 575:
V. Preliminary Evaluation of the Ec
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25-° 20/0M 15-a)CLU)0 10-0CL 5 -0)
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25 -- 20-0co/o 15--Da)a0a)10m~ C) 5
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forestlands. The costs of productio
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In Figure H.8, the equilibrium betw
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foregone and the way they will save
Page 586 and 587:
The income created by the purchase
Page 588 and 589:
Two concepts are useful for underst
Page 590 and 591:
y an estimate of the price that wou
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Table H.10. Comparison of timber em
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of the second year. The same assump
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are generally much lower than for c
Page 598 and 599:
582
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studies at University of Washington
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586
Page 604 and 605:
Most significant late successionall
Page 606 and 607:
to evaluate options. Information in
Page 608 and 609:
Q0Table 1.1. Estimated acres and ow
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soTable 1.3. Estimated acres and ow
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01CDarTable 1.5. Size class distrib
Page 614 and 615:
Table 1.8. Frequency distribution o
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Literature CitedThomas, J.W., E.D.
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602
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- The overlap of home ranges among
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Table J.2. Acreage and owl numbers
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continuedDCA Land AcreageIdent. Sta
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Page 628 and 629:
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Page 634 and 635:
Table J.9. Acrea ?e and owl numbers
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continued-DCA LandIdent. StatusAcre
Page 640 and 641:
-DCAIdent.LandStatusAcreageNRF Tota
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626
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Experience Includes: Assistant Dire
Page 646 and 647:
Education: B.S. in forestry from Or
Page 648 and 649:
Team SupportCharles Bruce, Oregon D
Page 650 and 651:
continued-1 5-24-91 Other SpeciesSu
Page 652 and 653:
continued-9-16 thru9-20-919-19-91Sp
Page 654 and 655:
Perry, D. 1991. The ecology of coar
Page 656 and 657:
2mandate the achievement of recover
Page 658 and 659:
642
Page 660 and 661:
644
Page 662 and 663:
Aspect - the direction a slope face
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Congressionally classified and desi
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Empirical - derived from direct obs
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Hummocky - a landscape characterize
Page 670 and 671:
Mixed conifer - as used in this doc
Page 672 and 673:
Potential habitat - a stand of tree
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Sensitive species - those species t
Page 676 and 677:
Take -Uunder the Endangered Species
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wild, scenic, or recreational by an
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