Recovery Plan for the Northern Spotted Owl - DRAFT

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Appendix CDemographic Analysis of NorthernSpotted Owl PopulationsDavid R. AndersonKenneth P. BurnhmnColorado Cooperative Fish and Wildlife Resource UnitColorado State University319
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I 1.2:Sp 6/draftRecovery Plan for t
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List of FiguresTable 4.1 Abbreviate
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Recovery ObjectiveSecretary of the
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Management Rules for Designated Con
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Monitoring and Research ProgramThe
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Chapter IIntroduction1
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"Threatened species" means a specie
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B. The Interagency Scientific Commi
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lished works and many government do
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II.A. Natural History of theNorther
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I- \IzXelT - - -.11 I..., ... I.."
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spring. Territories probably are sm
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Northern goshawks (Accipiter gentil
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also use a variety of nest sites in
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Home Range SizeHome range is define
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Table 2.2. Median amounts of old-gr
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3. Life Historycannot reject the hy
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Dispersal1984, Thomas et al. 1990,
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4. ConclusionOur knowledge of the n
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Ca n a d aEastern Washington Cascad
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Table 2.3. Estimated spotted owl ha
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Table 2.4. Significant threats to t
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Table 2.5. Results of surveys for s
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isolated. In these provinces, small
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centers for 30 to 35 spotted owl te
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Southwest Washington occupies a key
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Distribution of Habitat and Populat
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Province Isolation. The eastern Was
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were in suitable habitat condition.
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Estimated acres of forest landbase:
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owned and contain little suitable h
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Vulnerability to Natural Disturbanc
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Douglas-fir/hardwood types, the lat
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miles from the contiguous populatio
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In comparison, the southern part of
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Sacramento River Canyon now provide
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On January 11, 1980, there was an a
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vation Strategy for the Northern Sp
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transfer of one area to the Grand R
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The BLM Spokane (Washington) Distri
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3. National Park ServiceThe followi
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The results of a consultation are s
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- Since landowners already face cut
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4. Cumulative Effects AssessmentThe
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Assessment, Planning, and Monitorin
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Spotted owl nesting sites and activ
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Coastal Oregon Productivity Enhance
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including wildlife. In other cases,
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Currently, the Yakima Indian Nation
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where timber harvest is limited to
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III.A. Recovery Objective and Delis
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4. The population is unlikely to ne
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Needs of other species should be co
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years. The strategy of managing for
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't0
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Finally, the plan recommends mainta
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A total of 1,181 pairs of owls has
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DKnown owlsin the OregonprovincesKn
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mologists, and representatives of o
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this prey species is an appropriate
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After 100 years residual snags will
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2. Fuelwood Gathering. If allowed,
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The application of prescriptions A,
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owls. (Refer to tables in section I
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PRESCRIPTION D -RETAIN OWLS IN MANA
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126
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DCA Management Plans.The recovery p
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Endangered Species Act) or through
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6. Costs to landowners should be re
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The real or perceived disincentives
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authority of the public body to ent
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* Use the recovery plan's recommend
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140
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Supplemental pair areas - Habitat d
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Numbers of owls currently are estim
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Biological goals and implementation
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ning. Also, several unsurveyed area
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Owl pairsKnown owlsin the province3
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- Initiate long-range planning effo
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Table 3.8. Summary comments on the
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from nonfederal lands are needed to
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Eastern Washington Cascades Provinc
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(Owl pairs140 - _ _ _ _ _ _ _ _ _ _
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habitat in DCA WD- 16. Approximatel
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Table 3.12. Summary comments on the
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Ttllwrnook/Astoria area. Provide su
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connectivity between key DCAs. This
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ITable 3.14. Summary comments on th
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crest of the Cascade Mountains. The
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sites on federal lands (figure 3.21
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Owl pairsL I 350- _ _ . ...........
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DCAs, often combined with adjacent
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Owl pairsLiKnown owlsin the provinc
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nesting and roosting habitat until
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Option 3: Management of clusters: g
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Table 3.21. Summary of acreage and
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Owl pairs -Known owlsin the provinc
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and to within 0.5 miles of the acti
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Spotted owls have been found at 86
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their distance from other sites. Ar
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* Recommend population and habitat
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away in some or all of the range. D
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4. What are the population dynamics
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If the monitoring and research prog
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lowlands, eastern Oregon Cascades,
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210
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Stepdown Outline1. Management Tasks
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Table 4.1 Abbreviated Cost TableThe
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continued-TaskPriorityResp. PartyCo
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218
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220
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and the number of invertebrate anim
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Table 5.2. Numbers of other species
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Amphibians and Reptiles: Larch Moun
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228
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230
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Barrows, C. W. 1980. Feeding ecolog
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Clark, R. J., D. G. Smith, and L. H
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Franklin, A. B., J. A. Blakesley, a
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Harestad, A. S., and F. L. Bunnell.
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Lundquist, R. W. and J. M. Mariani.
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O'Halloran, K. 1989. Spotted owl in
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Solis, D. M. 1980. Habitat use by n
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Young, K. D., A. B. Franklin, and J
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248
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250
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Table A.1. General approaches for e
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complete count of the territorial b
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Table A.3. Powera for various desig
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Table A.4. Illustration of a Markov
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Obviously, these statements hold on
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Table A.6. Reliability of trend est
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Demographic AnalysisAs noted earlie
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the estimate (using the simple equa
Page 284 and 285: however, we know little about the b
Page 286 and 287: Table A.8. Summary of information n
Page 288 and 289: predict the power achieved by diffe
Page 290 and 291: Other studiestached. If dispersing
Page 292 and 293: impractical at present. We describe
Page 294 and 295: Conclusionsthe year effect can be i
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Page 302 and 303: owls. Consequently, the issue of wh
Page 304 and 305: Use of the informationAssessments a
Page 306 and 307: Recent LiteratureWe might begin by
Page 308 and 309: Kerms (1989) measured habitat struc
Page 310 and 311: Hamer (pers. comm.) described 11 ne
Page 312 and 313: Table B.3. Percent hardwoods in the
Page 314 and 315: Table B.5. Comparison of habitats u
Page 316 and 317: Table B.8. Tree density (number of
Page 318 and 319: and numbers of trees, and percent o
Page 320 and 321: Under this hypothesis, owl fitness
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Page 324 and 325: ResultsDiscussionspotted owl habita
Page 326 and 327: __Discussiondata from 41 sites in t
Page 328 and 329: Recommendations for Future Research
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Page 332 and 333: Hamer, T. 1988. Home range size of
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Page 338 and 339: Parameter Estimatesfor Individual S
Page 340 and 341: provided in Table C. 5. Study of th
Page 342 and 343: Table C.6. Summary of statistics re
Page 344 and 345: 3. ConclusionsThis source of bias i
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Page 348 and 349: Gray Wolf .........................
Page 350 and 351: formed during the initial meeting t
Page 352 and 353: provide habitat for the numerous sp
Page 354 and 355: The ApproachA committee of the Reco
Page 356 and 357: The List of Threatened and Endanger
Page 358 and 359: Mammalsoldest trees in a stand. Eig
Page 360 and 361: ABBFigure D. 1. Distribution of: a)
Page 362 and 363: KTable D.3. A list of threatened an
Page 364 and 365: Fishestheir dispersal capabilities
Page 366 and 367: OlympicNorthcoastalRockyMountainSou
Page 368 and 369: continued-StatusabState Considered
Page 370 and 371: continued-SpeciesColumbia sidebandM
Page 372 and 373: endemic to the Pacific Northwest).
Page 374 and 375: ------continued-SpeciesRhyacophila
Page 376 and 377: continued-SpeciesFed3tatusa bStateW
Page 378 and 379: StatusasbOld ForestState Associatio
Page 380 and 381: Summaryspores (Maser et al. 1978b).
Page 382 and 383: Biology of Owl Prey, Older Forest A
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Woodrats.-The dusky-footed woodrat
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and are less restrictive under stat
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Stock concept.-The term "stock" was
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Table D.10. List of fish stocks of
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continued-Steelhead-owipu Sea-runTr
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continued-BasinHoodWillametteRogueS
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Table D.13. Washington streams (or
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inland distribution currently decre
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nesting territories in the three st
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However, little is known about the
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Fisher (Martes penncmti)Distributio
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at one time supported marten popula
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each population of these animals is
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species also is found under the bar
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few meters from water after heavy r
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In considering the needs of other s
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Barrows, C. W. 1981. Roost selectio
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Clark, T. W., E. Anderson, C. Dougl
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Hamer, T. E. 1991. Habitat relation
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Mannan, R. W., E. C. Meslow, and H.
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Oakley, A. L., Collins, J. A., Ever
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Solis, D. M. 1983. Summer habitat e
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Washington Natural Heritage Program
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414
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f. Wilderness.g. Livestock grazing.
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I. Monitoring.1. Northern spotted o
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420
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here will reduce some of that habit
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424
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426
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Selection of Forest TypesAt the tim
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eceive as little as 25 inches. Disc
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A. FireThe combination and interact
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Table F.4. Important forest insects
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continued-Common Name Causal Agent
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D. DiseasesForest diseases in the P
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III. Forest Protection in theWest C
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120 100200 500I~~~~.K446Figure F.4.
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InsectsDiseasesInsects in this subr
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__hardwoods are immune. Shade toler
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pests (Schowalter 1988). Black stai
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tion and initial attack. If manipul
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log and snag density was likely low
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B. Management Effects on StandsMana
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units (underburning or lop/scatter)
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In the higher elevation White Fir a
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growth often increases (Reinhardt a
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50rcc4*0-cC)2M CmZ13010-yearI ,...*
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In the East Cascades subregion fore
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The introduced disease, white pine
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VI. Conclusionsmortality will resul
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Baker, F.A. 1988. The influence of
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Habeck, J.R. 1990. Old-growth ponde
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Perry, D.A., and J.G. Borchers. 199
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Weatherspoon, C.P., and C.N. Skinne
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482
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484
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area, structure, ages, location; ne
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Stand AFigure G.2. A--unthmnned Dou
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figure 0.3. Effects of thinning Dou
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II. Review of Natural Vegetation an
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western hemlock, and sometimes othe
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year intervals (Appendix F). With t
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Live crown ratiobecent)6050IStand A
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Basal area growth continued to incr
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technology probably will allow some
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Stand A. No activities (west side 3
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Stand A. No activities (west side o
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acre, 4 to 12 inches in diameter),
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allow redwood sprout clumps maximum
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KAge 15 years - 511 trees per acre,
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6Stems per acre100 _ Stand A. Natur
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Basal Area(square feetper acre)50 1
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* Opening the canopy in small patch
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Drew, T.J. and J.W. Flewelling. 197
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Miller, R.E.; D.L. Reukema, and R.L
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Tappeiner, J.C., P.M. McDonald, and
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526
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528
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6. Timber supply curve and economic
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scientifically credible biological
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E. Summary of Preliminary Estimates
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II. General Economic Principles for
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practices may be able to show how s
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Economic patterns show the efficien
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The sustainable yield concept that
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A. Previously Estimated Economic Co
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County receipts were estimated to d
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B. Potential Social Costs of Owl Co
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The contrast between the behavior d
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0U1 n1nDTable H.2. Economic depende
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cḻlTable H.4. Economic characteri
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An analysis completed by the State
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V. Preliminary Evaluation of the Ec
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25-° 20/0M 15-a)CLU)0 10-0CL 5 -0)
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25 -- 20-0co/o 15--Da)a0a)10m~ C) 5
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forestlands. The costs of productio
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In Figure H.8, the equilibrium betw
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foregone and the way they will save
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The income created by the purchase
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Two concepts are useful for underst
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y an estimate of the price that wou
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Table H.10. Comparison of timber em
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of the second year. The same assump
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are generally much lower than for c
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582
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studies at University of Washington
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586
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Most significant late successionall
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to evaluate options. Information in
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Q0Table 1.1. Estimated acres and ow
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soTable 1.3. Estimated acres and ow
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01CDarTable 1.5. Size class distrib
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Table 1.8. Frequency distribution o
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Literature CitedThomas, J.W., E.D.
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602
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- The overlap of home ranges among
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Table J.2. Acreage and owl numbers
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continuedDCA Land AcreageIdent. Sta
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Table J.9. Acrea ?e and owl numbers
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continued-DCA LandIdent. StatusAcre
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-DCAIdent.LandStatusAcreageNRF Tota
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626
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Experience Includes: Assistant Dire
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Education: B.S. in forestry from Or
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Team SupportCharles Bruce, Oregon D
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continued-1 5-24-91 Other SpeciesSu
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continued-9-16 thru9-20-919-19-91Sp
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Perry, D. 1991. The ecology of coar
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2mandate the achievement of recover
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642
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644
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Aspect - the direction a slope face
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Congressionally classified and desi
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Empirical - derived from direct obs
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Hummocky - a landscape characterize
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Mixed conifer - as used in this doc
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Potential habitat - a stand of tree
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Sensitive species - those species t
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Take -Uunder the Endangered Species
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wild, scenic, or recreational by an
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