Recovery Plan for the Northern Spotted Owl - DRAFT

Recovery Plan for the Northern Spotted Owl - DRAFT Recovery Plan for the Northern Spotted Owl - DRAFT

10.07.2015 Views

Recent LiteratureWe might begin by examining the data from utilized habitats. Finding thatutilized habitats frequently met our definition of Type A habitat, and that homeranges consistently contained large amounts of Type A habitat, would supportthe hypothesis. We would also examine data from correlational analyses. Thehabitat categories would be Type A and other. Strong preference for Type Astands would tend to support our hypothesis, as would a finding that densityand demographic rates were higher in circles, landscapes, and territories withhigh amounts of Type A habitat. Results from functional studies would beuseful in determining whether the specific features of importance to owls werehighly correlated with the elements in our definition of Type A habitat and ifthey were, whether this was likely to be true in other environments to which wemight wish to extrapolate our findings. We would not expect that all these testsand analyses to support the predictions, but if few of them did, then we probablywould discard or substantially modify the definition of Type A habitat andstart again. If most of the predictions were verified, then we would tend to feelthat a good definition of suitable habitat had been developed.A process very much like that above has been carried out in evaluating thehypothesis that older (i.e., mature and old-growth) forests provide suitablehabitat for owls. Descriptions of utilized habitat have shown that nesting,roosting, and foraging owls usually are found in older stands and the structuralfeatures of these stands has been described in quantitative terms. Comparisonof stands have shown that owls exhibit a strong preference for olderstands for roosting and foraging. Reproductive success has been shown to behigher for pairs having more older forest in or near their home ranges. Densityand adult turnover rates also have been shown to be positively correlated withamount of older forest in the landscape. Several hypotheses about the possiblefunctions of older forest in supporting owls have been generated, and evidencesupporting some of them has been reported. Thus, while no one piece ofevidence would (or should) provide high confidence that older forest providessuitable habitat for owl populations, taken together these studies do providesuch confidence. This of course does not mean that older forest is the onlyhabitat that provides suitable habitat, but hypotheses predicting that otherhabitats also provide suitable habitat must be evaluated in the same comprehensiveway that the hypothesis about older forest has been evaluated.As the previous examples indicate, the results from descriptive, correlational,and functional studies are best used in combination, rather than in isolation.Descriptive information is most useful for defining habitat categories; correlationalinformation at the stand, home range, and landscape level is most usefulfor measuring suitability of specific habitats; functional information is particularlyuseful for identifying the specific resources of value to owls and thusproviding insights about how widely results can be extrapolated to otherpopulations. Information from any single type of study would provide a poorbasis for assessments or predictions about habitat suitability: taken together,however, they provide an excellent basis for such analyses. Subsequentsections explore in more detail the ways that each type of information contributesto analyses of habitat suitability.Thomas et al. (1990) provided a thorough review of the literature, includingmany pre-publication reports, available at the time of their report. Our discussionof habitat suitability is based largely on their review. Several reports haveappeared since their study, and we have conducted new analyses of a fewtopics. The new studies, or studies from which we have extracted originaldata, are described briefly in the following section.290

CaliforniaAsrow (1983) sampled the vegetation of four occupied spotted owl managementareas (SOMAs) in old-growth Douglas-fir/white fir forests on the Scott RiverRanger District of the Klamath National Forest. Owl presence was documentedusing a combination of night and day surveys during the 1983 breedingseason. Vegetation was sampled on six variable-radius plots in each SOMA.One plot was centered on the location of the bird; other plots were placed 132feet from the central plot and equal distances from each other.Bingham (1991) described stands used by 20 radio-transmittered owls forroosting and foraging in Douglas-fir/tanoak/madrone forests on the Mad RiverRanger District (Six Rivers National Forest) in the Klamath province. Utilizedstands were defined as those with more than 3 telemetry locations. Data weresummarized separately for roosting stands (based on daytime locations) andforaging stands (based on nighttime locations) in the breeding and nonbreedingseason. Vegetation was sampled on 3 to 8 0.25-acre plots per stand. Theauthor presented the data as means for each of three vegetation classes(defined on the basis of a cluster analysis): the data presented in this appendixare' the means of these three classes, weighted by sample sizes.Blakesley et al. (1992) recorded 421 owl locations, including 79 nest sites,during the breeding seasons of 1985 to 1989 in the Willow Creek study area innorthwest California. Stand types were defined by dominant vegetation (conifers,hardwoods, unvegetated), mean dbh, elevation, aspect, position (lower,middle, or upper third), and slope.Chavez-Leon (1989) measured habitat structure in 14 stands used by owlsduring 1987 and 1988 in the Klamath province (Humboldt and MendocinoCounties) in northwestern California. Pairs occupied nine and 10 sites in 1987and 1988, respectively: nesting pairs occupied five and three sites in 1987 and1988, respectively. Twelve of the 14 sites were in typical Douglas-fir/tanoak/madrone forests, and two were in redwoods. The landscape was highly fragmented:stands used by owls sometimes were fragmented and varied in sizefrom 49 to 642 acres. Habitat structure was measured on five or more variable-radiusplots randomly placed within each stand. Chavez-Leon excludedsome plots in calculating mean densities and basal areas, but we used all plotsfor our analysis.Folliard and Reese (1991) measured 30 nest sites and nest stands during 1990in second-growth redwood/Douglas-fir forests of the California Coast province(Humboldt and Del Norte Counties). Plots were circular and covered 0.18 acres(radius = 50 feet). Nest-site plots were centered on the nest but did not includethe nest tree in measures of density and basal area. Four or five plots weredistributed randomly within 0.5 miles of the nest and in the nest stand.Jimerson et al. (1991a) quantified the structure of old-growth Douglas-fir/tanoak/madrone stands in SAF 234 (i.e., Society of American Foresters foresttype 234) in the Klamath province. Data were collected on the Six Rivers,Siskiyou, and Klamath (western half National Forests, and the NorthernCalifornia Coast Range Preserve. Vegetation was sampled on three variableradiusplots in each old-growth stand. Bingham and Sawyer (1991) presenteda summary of this information.Jimerson et al. (1991b) quantified the structure of old-growth Pacific Douglasfirforests stands in SAF 229 in the Klamath province. Data were collected onthe western half of the Klamath and Six Rivers National Forests. Vegetationwas sampled on three variable-radius plots in each old-growth stand.291

Cali<strong>for</strong>niaAsrow (1983) sampled <strong>the</strong> vegetation of four occupied spotted owl managementareas (SOMAs) in old-growth Douglas-fir/white fir <strong>for</strong>ests on <strong>the</strong> Scott RiverRanger District of <strong>the</strong> Klamath National Forest. <strong>Owl</strong> presence was documentedusing a combination of night and day surveys during <strong>the</strong> 1983 breedingseason. Vegetation was sampled on six variable-radius plots in each SOMA.One plot was centered on <strong>the</strong> location of <strong>the</strong> bird; o<strong>the</strong>r plots were placed 132feet from <strong>the</strong> central plot and equal distances from each o<strong>the</strong>r.Bingham (1991) described stands used by 20 radio-transmittered owls <strong>for</strong>roosting and <strong>for</strong>aging in Douglas-fir/tanoak/madrone <strong>for</strong>ests on <strong>the</strong> Mad RiverRanger District (Six Rivers National Forest) in <strong>the</strong> Klamath province. Utilizedstands were defined as those with more than 3 telemetry locations. Data weresummarized separately <strong>for</strong> roosting stands (based on daytime locations) and<strong>for</strong>aging stands (based on nighttime locations) in <strong>the</strong> breeding and nonbreedingseason. Vegetation was sampled on 3 to 8 0.25-acre plots per stand. Theauthor presented <strong>the</strong> data as means <strong>for</strong> each of three vegetation classes(defined on <strong>the</strong> basis of a cluster analysis): <strong>the</strong> data presented in this appendixare' <strong>the</strong> means of <strong>the</strong>se three classes, weighted by sample sizes.Blakesley et al. (1992) recorded 421 owl locations, including 79 nest sites,during <strong>the</strong> breeding seasons of 1985 to 1989 in <strong>the</strong> Willow Creek study area innorthwest Cali<strong>for</strong>nia. Stand types were defined by dominant vegetation (conifers,hardwoods, unvegetated), mean dbh, elevation, aspect, position (lower,middle, or upper third), and slope.Chavez-Leon (1989) measured habitat structure in 14 stands used by owlsduring 1987 and 1988 in <strong>the</strong> Klamath province (Humboldt and MendocinoCounties) in northwestern Cali<strong>for</strong>nia. Pairs occupied nine and 10 sites in 1987and 1988, respectively: nesting pairs occupied five and three sites in 1987 and1988, respectively. Twelve of <strong>the</strong> 14 sites were in typical Douglas-fir/tanoak/madrone <strong>for</strong>ests, and two were in redwoods. The landscape was highly fragmented:stands used by owls sometimes were fragmented and varied in sizefrom 49 to 642 acres. Habitat structure was measured on five or more variable-radiusplots randomly placed within each stand. Chavez-Leon excludedsome plots in calculating mean densities and basal areas, but we used all plots<strong>for</strong> our analysis.Folliard and Reese (1991) measured 30 nest sites and nest stands during 1990in second-growth redwood/Douglas-fir <strong>for</strong>ests of <strong>the</strong> Cali<strong>for</strong>nia Coast province(Humboldt and Del Norte Counties). Plots were circular and covered 0.18 acres(radius = 50 feet). Nest-site plots were centered on <strong>the</strong> nest but did not include<strong>the</strong> nest tree in measures of density and basal area. Four or five plots weredistributed randomly within 0.5 miles of <strong>the</strong> nest and in <strong>the</strong> nest stand.Jimerson et al. (1991a) quantified <strong>the</strong> structure of old-growth Douglas-fir/tanoak/madrone stands in SAF 234 (i.e., Society of American Foresters <strong>for</strong>esttype 234) in <strong>the</strong> Klamath province. Data were collected on <strong>the</strong> Six Rivers,Siskiyou, and Klamath (western half National Forests, and <strong>the</strong> Nor<strong>the</strong>rnCali<strong>for</strong>nia Coast Range Preserve. Vegetation was sampled on three variableradiusplots in each old-growth stand. Bingham and Sawyer (1991) presenteda summary of this in<strong>for</strong>mation.Jimerson et al. (1991b) quantified <strong>the</strong> structure of old-growth Pacific Douglasfir<strong>for</strong>ests stands in SAF 229 in <strong>the</strong> Klamath province. Data were collected on<strong>the</strong> western half of <strong>the</strong> Klamath and Six Rivers National Forests. Vegetationwas sampled on three variable-radius plots in each old-growth stand.291

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