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Crop yield response to water - Cra

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Figure 9Wine quality score as a function of <strong>to</strong>tal irrigation for Cabernet Sauvignon in Adulam, Israel.Source: Bravdo et al. (1985).20 bunches per vine 40 bunches per vine 60 bunches per vine1413Wine quality score1211109200 250 300 350 400 450Irrigation (mm)pruning, thinning, and canopy management maybe important <strong>to</strong> achieve both high <strong>yield</strong> andhigh quality.Water deficit generally increases the concentration of phenolic compounds, but has a differentialeffect on individual groups of phenols depending on timing and severity of the stress. Tanninbiosynthesis can be negatively affected by severe stress after anthesis (Ojeda, 2002), but laterdeficits often increase tannin concentration (Roby et al., 2004). In most cases, anthocyaninconcentration responds positively <strong>to</strong> <strong>water</strong> shortage after veraison but less frequently <strong>to</strong>pre-veraison deficits (Matthews and Anderson, 1988); although the expression of genes involvedin anthocyanin biosynthesis can be increased by <strong>water</strong> stress pre-veraison (Castellarin et al.,2007). Aside from phenolic compounds, aroma attributes may also be affected (Chapman et al.,2005). Differences in the <strong>response</strong> of varieties are likely but not well documented (see: SuggestedRDI regimes below for examples). In addition <strong>to</strong> the effects on amount and proportion of keycompounds, <strong>water</strong> deficit can cause more subtle but relevant effects. For example, <strong>water</strong> deficitafter veraison has been shown <strong>to</strong> increase the structural complexity (degree of polymerisation)and <strong>to</strong> reduce the extractability of phenolic compounds in berries of several red varieties (Ojedaet al., 2002; Sivilotti et al., 2005). The effects of <strong>water</strong> deficit on berry attributes are partiallyrelated <strong>to</strong> reductions in berry size, although size-independent effects have also been reported.Allometric analysis is required (i) <strong>to</strong> separate size-dependent effects of <strong>water</strong> deficit on aparticular component, e.g. sugar, and (ii) <strong>to</strong> compare the relative responsiveness of differentberry components <strong>to</strong> <strong>water</strong> deficit (Sadras et al., 2007 and Sadras and McCarthy, 2007). Forexample, allometric analysis revealed that <strong>water</strong> deficit accelerated the rate of accumulation ofanthocyanins with respect <strong>to</strong> sugar of Cabernet Sauvignon in a warm environment (Figure 12).Water management is therefore important <strong>to</strong> ensure a certain coupling of key berry componentsduring ripening; this will eventually affect wine balance.472crop <strong>yield</strong> <strong>response</strong> <strong>to</strong> <strong>water</strong>

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