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Crop yield response to water - Cra

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Early vegetative and reproductive growthUnder subtropical climate conditions in the Northern Hemisphere (California, for example),the main vegetative growth flush occurs in late February and March. There are normallyadditional growth flushes in the summer and autumn. Leaves are viable for about two years.Leaves are continuously replenished although leaf drop is generally highest during the spring.Most citrus cultivars produce flowers in the spring and are self-compatible; they may befertilized by their own pollen. In cool, coastal climates, flowers may be produced throughoutthe year but maximum flowering is also in the spring. Flowers develop in leaf axils on oldershoots as opposed <strong>to</strong> shoots developing in growth flushes.Citrus trees produce a large amount of flowers resulting in small fruit. However, a largepercentage of these small fruit drop from the tree apparently because of a combination ofphysiological and environmental fac<strong>to</strong>rs. There appears <strong>to</strong> be a hierarchy of flowers relative<strong>to</strong> fruit drop; flower in locations with a higher flower/leaf ratio are more subject <strong>to</strong> drop.Flowers that open late in the bloom period are more likely <strong>to</strong> set fruit that survives <strong>to</strong> harvestthan fruit set from early flowers. Likewise, faster growing fruit is less apt <strong>to</strong> drop than slowgrowing fruit.The primary fac<strong>to</strong>r controlling the dropping of flowers and young fruit is a weakening oftissue in a preformed abscission zone at the point of attachment at the base of the ovary<strong>to</strong> the disk or of the pedicel <strong>to</strong> the stem. The actual mechanism of the process is not wellunders<strong>to</strong>od but is believed <strong>to</strong> be triggered by growth regula<strong>to</strong>rs in <strong>response</strong> <strong>to</strong> either externalor internal changes. Young fruit that remain after the first period of dropping presumably arecapable of developing <strong>to</strong> maturity if the weather is favourable. Nevertheless, some droppingof fruit occurs throughout the season. As the weather becomes hotter in the early summermonths, there is generally a period of accelerated fruit drop, which is often referred <strong>to</strong> asthe ‘June drop’ period. If the heat is severe and prolonged for several days, fruit drop can beheavy, resulting in a greatly reduced crop. This has been observed in several cultivars of thenavel and Valencia types. Some cultivars, notably Valencia and mandarins, have an alternatebearing cycle; light crop years following heavy crop years.Fruit development stageFollowing early season fruit drop, and after the leaves of new flushes have fully expanded, theremaining fruit begin <strong>to</strong> develop initially by rapid cell division. At this point, a cross-section ofthe fruit clearly reveals its component parts:• Flavedo (epicarp) – a rough, robust and bright colour (from yellow <strong>to</strong> orange) skin or rindthat covers the fruit and protects it from damage. Its glands contain the essential oils thatgive the fruit its typical citrus fragrance.• Albedo (mesocarp) – a white, spongy tissue that, <strong>to</strong>gether with the flavedo, forms the peel(pericarp) of the fruit. • Pulp (endocarp) – the internal part is comprised of individual segments or juice sacs. This isthe edible part of the fruit and the source of the juice.318crop <strong>yield</strong> <strong>response</strong> <strong>to</strong> <strong>water</strong>

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