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Crop yield response to water - Cra

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ox 2 Understanding the transpiration process.Water loss from plant leaves takes place primarily through pores called s<strong>to</strong>mata thatare opened during daylight hours and close at night. S<strong>to</strong>mata are also the path forthe entrance of carbon dioxide in<strong>to</strong> the leaves, the necessary input for pho<strong>to</strong>synthesis.Transpiration losses from leaves trigger a sequence of events in the soil tree system;<strong>water</strong> moves from the shoots in<strong>to</strong> the leaves <strong>to</strong> compensate for leaf losses, drawing<strong>water</strong> from the trunk in<strong>to</strong> the shoots; this in turn, draws <strong>water</strong> from the roots <strong>to</strong> thetrunk, and finally, from the soil <strong>to</strong> the roots. Thus, liquid <strong>water</strong> flows from the soil<strong>to</strong> the sites in the leaves where it evaporates before diffusing through the s<strong>to</strong>mata<strong>to</strong> the atmosphere. Water moves passively through this path following a gradientin potential energy, from the soil <strong>to</strong> the leaves. In any location along this path, suchenergy level may be determined and is called <strong>water</strong> potential. Furthermore, the <strong>water</strong>flow encounters resistance all along the pathway from the soil <strong>to</strong> the atmosphere (seediagram). The tree exerts some control on this <strong>water</strong> flow, which is needed <strong>to</strong> matchthe transpirational loss <strong>to</strong> the evaporative demand of the environment. When there areimbalances between supply and demand, the tree controls its <strong>water</strong> loss through theadjustment of the degree of s<strong>to</strong>matal opening and by other means.The <strong>water</strong> flow from soil through tree <strong>to</strong> the atmosphere.Yield Response <strong>to</strong> Water of Fruit Trees and Vines 255

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