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Crop yield response to water - Cra

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size as affected by soil <strong>water</strong> deficit, and s<strong>to</strong>matal control plays a minor role. After anthesis,the leaves are fully grown and consequently the control of transpiration is more dependentupon s<strong>to</strong>matal closure and the extent of canopy senescence. Periods of <strong>water</strong> deficit at anygrowth stage can cause canopy senescence with subsequent reduction in seed <strong>yield</strong>. Thereis genetic variability in the <strong>response</strong> of sunflower genotypes <strong>to</strong> <strong>water</strong> deficits. Long-seasongenotypes have greater canopy cover and produce more biomass under drought conditions,because of their ability <strong>to</strong> extract more <strong>water</strong> from the subsoil (Gimenez and Fereres, 1986).Also, it has been observed that sunflower cultivars adapted <strong>to</strong> an arid climate are less sensitive<strong>to</strong> <strong>water</strong> stress than the cultivars developed for humid climates.Under <strong>water</strong> stress, the time between planting and flowering remains relatively constant, andinflorescence initiation is also relatively insensitive <strong>to</strong> <strong>water</strong> stress. Subsequent developmen<strong>to</strong>f the inflorescence is, however, affected by <strong>water</strong> deficits and <strong>water</strong> stress reduces thenumber of flowers. The entire flowering period is thus the most sensitive <strong>to</strong> <strong>water</strong> deficits asthe number of seeds may be negatively affected. Seed filling following flowering is the nextmost sensitive period <strong>to</strong> <strong>water</strong> deficits that reduce both seed weight, seed number (because ofabortion) and oil content. Therefore, the reproductive stages (flowering and ripening stages)are more sensitive <strong>to</strong> <strong>water</strong> stress than the vegetative stages. Maintenance of green leaf areaand pho<strong>to</strong>synthesis after anthesis is key, as seed weight and oil content are mostly influencedby intercepted radiation and carbon assimilation during seed filling. During this phase, boths<strong>to</strong>matal closure and leaf senescence play a role in the control of plant <strong>water</strong> status. On theother hand, there is the associated cost of reduced CO 2 assimilation. The contribution thatcarbon fixed before anthesis makes <strong>to</strong> grain-filling of sunflower may be important under<strong>water</strong> stress after anthesis. Nevertheless, the harvest index of sunflower is often negativelyaffected by <strong>water</strong> deficits during the reproductive phase.Genotypes having a gradual <strong>response</strong> <strong>to</strong> <strong>water</strong> stress may be most suited <strong>to</strong> environmentswith severe <strong>water</strong> deficits. In environments with short, frequent and moderate soil <strong>water</strong>deficits alternating with well-<strong>water</strong>ed periods, maintaining organ expansion and biomassproduction would result in better agronomic performance. Several studies of sunflower havedemonstrated that there is a close relationship between canopy size at flowering and cropseed <strong>yield</strong>. The plasticity of the crop, in terms of adapting leaf area development <strong>to</strong> <strong>water</strong>availability, is well known.Sunflower is moderately <strong>to</strong>lerant <strong>to</strong> salinity, being unaffected by soil salinity up <strong>to</strong> 4.8 dS/m. Itis currently cultivated in dry areas where salinity can be a threat. Thus, it could be adapted <strong>to</strong>salt-affected soils, provided irrigation management is adequate and salt leaching is practised.In all cases, leaching is important <strong>to</strong> limit salt accumulation in the root zone, avoiding thereduction of seed and oil <strong>yield</strong> under salt stress. It would be feasible <strong>to</strong> use moderately saline<strong>water</strong> <strong>to</strong> irrigate sunflower. Nevertheless it is necessary <strong>to</strong> maintain high soil <strong>water</strong> content inthe root zone all the time, and <strong>to</strong> minimize salinity built up by leaching.The nutrient content extracted by a sunflower crop producing around 1 <strong>to</strong>nne of seed <strong>yield</strong>per hectare includes around: 50 kg/ha of N, 15 kg/ha of P 2 O 5 , and 35 kg/ha of K 2 O. Fertilizerapplication depends on the expected <strong>yield</strong>s and the residual nutrients, varying from 20 <strong>to</strong> 140kg/ha of N, 15 <strong>to</strong> 70 kg/ha of P 2 O 5 , and 15 <strong>to</strong> 150 kg/ha of K 2 O.168crop <strong>yield</strong> <strong>response</strong> <strong>to</strong> <strong>water</strong>

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