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Crop yield response to water - Cra

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minimize <strong>water</strong> deficits during the three periods already mentioned: (a) tillering <strong>to</strong> stemelongation, (b) time of flowering, and (c) early <strong>to</strong> mid-grain filling. After the development of areasonable canopy, subsequent irrigations may be scheduled such that up <strong>to</strong> 50-60 percent of<strong>to</strong>tal available soil <strong>water</strong> (TAW) is depleted between applications without a notable negativeimpact on <strong>yield</strong> (Geerts and Raes, 2009).Over irrigation is common in wheat, even under supplemental irrigation: two or threeirrigations are sometimes applied in a short interval with little consideration of soil <strong>water</strong>status or crop demand. Excessive <strong>water</strong> supply results in lower <strong>water</strong> productivity in termsof <strong>yield</strong> per unit of <strong>water</strong> applied. In extreme situations, excessive <strong>water</strong> (from rainfall orirrigation) results in <strong>water</strong>logging which, as mentioned earlier, can substantially depressgrowth and <strong>yield</strong>. Over-generous <strong>water</strong> supply during the vegetative period, particularly incombination with high fertility, produces luxuriant vegetation and may result in lodging afterhead formation. Lodging may also occur if an excessive amount of <strong>water</strong> is applied in a singleirrigation late in development, particularly with sprinkler irrigation.YieldWheat grain <strong>yield</strong>s (at 11 percent moisture) can vary from crop failures in seasons with less than 100<strong>to</strong> 150 mm available <strong>water</strong>, <strong>to</strong> 1-3 <strong>to</strong>nne/ha in <strong>water</strong> limited rainfed conditions (Mediterranean,arid, dry-season subtropical environments) and 4-10 <strong>to</strong>nne/ha in rainfed temperate (westernand northern Europe) climates or irrigated systems (China). Exceptionally, grain <strong>yield</strong>s can reacha maximum of 15 <strong>to</strong>nne/ha in cool, long season (life cycle of over 300 days) environments withhigh solar radiation input such as southern New Zealand, Southern Chile, Ireland, England andsome regions of China. In 2009, average country <strong>yield</strong>s ranged from less than 0.5 <strong>to</strong>nne/ha inHonduras, Lesotho, Somalia, Venezuela and Eritrea <strong>to</strong> more than 9 <strong>to</strong>nne/ha in Belgium.A major fac<strong>to</strong>r contributing <strong>to</strong> the improvement in <strong>yield</strong> over the last century is the increases ofHI brought about by breeding for shorter statue. Under favourable conditions with no stress,HI ranges between 0.45 and 0.55 for modern wheat cultivars (Austin, 1999). However, whenthere is <strong>water</strong> stress after flowering or when the cultivar is poorly matched <strong>to</strong> the productionenvironment, HI can fall <strong>to</strong> as low as 0.20 <strong>to</strong> 0.30.The balance of <strong>water</strong> supply before and after flowering can have a substantial effect on grainquality. Water stress during grain filling leads <strong>to</strong> shrivelled grain with a low milling percentage(flour produced per grain input). On the other hand, high <strong>water</strong> supply late in the seasonleads <strong>to</strong> increased <strong>yield</strong> with low protein concentration. These changes alter the suitability ofthe grain for various end uses.wheat 99

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