Composite Training Unit Exercises and Joint Task ... - Govsupport.us
Composite Training Unit Exercises and Joint Task ... - Govsupport.us Composite Training Unit Exercises and Joint Task ... - Govsupport.us
COMPTUEX/JTFEX EA/OEA Final Chapter 3California or adjacent waters, although sightings have become more frequent in the past 20 years. Theminimum population estimate of the California/Oregon/Washington Stock is 12,748 (CV=0.28), based onship surveys conducted between 1991 and 1996 (Carretta et al., 2005).Distribution—A comprehensive study of the distribution of Risso’s dolphin in the Gulf of Mexico foundthat they used the steeper sections of the upper continental slope in waters 1,150–3,200 ft (350–975 m)deep (Baumgartner, 1997). Risso’s dolphins have been sighted in waters of the SOCAL Range Complexduring all seasons. However, in most years, higher numbers are present during the cold-water monthsthan during other times of the year (Forney and Barlow, 1998). Most sightings in the study area havebeen well offshore, but Risso’s dolphins have been sighted close to the eastern shore of San ClementeIsland during the cold season (Carretta et al., 2000). Risso’s dolphins occur individually or in small tomoderate-sized groups, normally ranging in numbers from 2 to nearly 250. The majority of groupscontain fewer than 50 (Leatherwood et al., 1980; Carretta et al., 1995 and 2000), however group sizesmay reach as high as 2,500. Risso’s dolphins are found in the SOCAL OPAREA throughout the year(Carretta et al., 2000).Acoustics—Risso’s dolphin vocalizations include broadband clicks, barks, buzzes, grunts, chirps,whistles, and simultaneous whistle and burst-pulse sounds (Corkeron and Van Parijs, 2001). Thecombined whistle and burst pulse sound appears to be unique to Risso’s dolphin (Corkeron and VanParijs, 2001). Corkeron and Van Parijs (2001) recorded five different whistle types, ranging in frequencyfrom 4 to 22 kHz. Broadband clicks had a frequency range of 6 to greater than 22 kHz. Low-frequencynarrowband grunt vocalizations had a frequency range of 0.4 to 0.8 kHz. A recent study establishedempirically that Risso’s dolphins echolocate; estimated source levels were up to 216 dB re 1 Pa-m(Philips et al., 2003).Nachtigall et al. (1995) conducted baseline audiometric work. Because of the natural background noise(the study was conducted in a natural setting), it was not possible to precisely determine peak (or best)hearing sensitivity in the species. Maximum sensitivity occurred between 8 and 64 kHz. Reportedthresholds (all re 1 Pa-m) were 124 dB at 1.6 kHz, 71.7 dB at 4 kHz, 63.7 dB at 8 kHz, 63.3 dB at 16kHz, 66.5 dB at 32 kHz, 67.3 dB at 64 kHz, 74.3 dB at 80 kHz, 124.2 dB at 100 kHz, and 122.9 dB at 110kHz.Rough-toothed Dolphin (Steno bredanensis)Status—The rough-toothed dolphin is not listed as endangered under the ESA or as depleted or strategicunder the MMPA. There are no abundance estimates available for this species in the NOAA StockAssessment Report for this area of the Pacific.Distribution—Rough-toothed dolphins are typically found in tropical and warm temperate waters (Perrinand Walker, 1975 in Bonnell and Dailey, 1993), rarely ranging north of 40°N or south of 35°S (Miyazakiand Perrin, 1994). Rough-toothed dolphins occur in low densities throughout the ETP where surfacewater temperatures are generally above 25°C (Perrin and Walker, 1975). Sighting and stranding recordsin the eastern North Pacific Ocean are rare (e.g., Ferrero et al., 1994).Rough-toothed dolphins usually form groups of 10–20 (Reeves et al., 2002), but aggregations of hundredscan be found (Leatherwood and Reeves, 1983). In the ETP, they have been found in mixed groups withspotted, spinner, and bottlenose dolphins (Perrin and Walker, 1975). Reeves et al. (2002) suggested thatthey are deep divers, and can dive for up to 15 min. They usually inhabit deep waters (Davis et al., 1998),where they prey on fish and cephalopods (Reeves et al., 2002). There have been few sightings of rough-February 2007 3-52
COMPTUEX/JTFEX EA/OEA Final Chapter 3toothed dolphins in the SOCAL OPAREA; therefore seasonal occurrence can not be determined (Ferreroet al., 1994).Acoustics—The vocal repertoire of the rough-toothed dolphin includes broad-band clicks, barks, andwhistles (Yu et al. 2003). Echolocation clicks of rough-toothed dolphins are in the frequency range of 0.1to 200 kHz, with a peak of about 25 kHz (Miyazaki and Perrin, 1994; Yu et al., 2003). Whistles show awide frequency range: 0.3 to >24 kHz (Yu et al., 2003). There is no published information on hearingability of this species.Short-beaked Common Dolphin (Delphinus delphis)Status—This is the most abundant cetacean off California (Dohl et al., 1981; Forney et al., 1995; Carrettaet al., 2005). The single current management unit for the short-beaked common dolphin in this area is aCalifornia/Oregon/Washington Stock, with a minimum population estimate of 449,846 (CV = 0.25)individuals (Carretta et al. 2004). The abundance of common dolphins varies seasonally but may beincreasing in California with a northward shift in the population (Heyning and Perrin, 1994; Barlow et al.,1997; Forney, 1997). The short-beaked common dolphin is not listed as endangered under the ESA or asdepleted or strategic under the MMPA.Distribution—Along the U.S. west coast, the short-beaked common dolphins’ distribution overlaps withthat of the long-beaked common dolphin. The short-beaked common dolphin is distributed between thecoast and at least 556 km from shore (Carretta et al., 2005). Short-beaked common dolphin abundanceoff California has increased dramatically since the late 1970s, along with a concomitant decrease inabundance in the ETP, suggesting a large-scale shift in the distribution of this species in the eastern NorthPacific (Forney et al., 1995; Forney and Barlow, 1998). The northward extent of short-beaked commondolphin distribution appears to vary inter-annually and with changing oceanographic conditions (Forneyand Barlow 1998). Short-beaked common dolphins are found in the SOCAL OPAREA throughout theyear (Forney and Barlow, 1998).Stomach contents of Delphinus from California waters revealed 19 species of fish and 2 species ofcephalopods; Delphinus feeds primarily on organisms in the vertically migrating DSL (Evans, 1994).Diel fluctuations in vocal activity of this species (more vocal activity during late evening and earlymorning) appear to be linked to feeding on the DSL as it rises during the same time (Goold, 2000). Atagged individual tracked off San Diego conducted dives deeper than 200 m, but with most in the range of9 to 50 m (Evans, 1971, 1994).Acoustics—Recorded Delphinus vocalizations include whistles, chirps, barks, and clicks (Ketten, 1998).Clicks and whistles have dominant frequency ranges of 23 to 67 kHz and 0.5 to 18 kHz, respectively(Ketten, 1998). Maximum source levels were approximately 180 dB 1 Pa-m (Fish and Turl, 1976).Oswald et al. (2003) found that short-beaked common dolphins in the ETP have whistles with a meanfrequency range of 6.3 kHz, mean maximum frequency of 13.6 kHz, and mean duration of 0.8 sec. Popovand Klishin (1998) recorded auditory brainstem responses from a common dolphin. The audiogram wasU-shaped with a steeper high-frequency branch. The audiogram bandwidth was up to 128 kHz at a levelof 100 dB above the minimum threshold. The minimum thresholds were observed at frequencies of 60 to70 kHz.Short-finned Pilot Whale (Globicephala macrorhynchus)Status—The short-finned pilot whale is not listed under the ESA. However, theCalifornia/Oregon/Washington Stock is considered strategic under the MMPA because the averagehuman-caused mortality may not be sustainable (Barlow et al., 1997). A recent calculation of the3-53 February 2007
- Page 65 and 66: COMPTUEX/JTFEX EA/OEA Final Chapter
- Page 67 and 68: COMPTUEX/JTFEX EA/OEA Final Chapter
- Page 69 and 70: COMPTUEX/JTFEX EA/OEA Final Chapter
- Page 71 and 72: COMPTUEX/JTFEX EA/OEA Final Chapter
- Page 73 and 74: COMPTUEX/JTFEX EA/OEA Final Chapter
- Page 75 and 76: COMPTUEX/JTFEX EA/OEA Final Chapter
- Page 77 and 78: COMPTUEX/JTFEX EA/OEA Final Chapter
- Page 79 and 80: COMPTUEX/JTFEX EA/OEA Final Chapter
- Page 81 and 82: COMPTUEX/JTFEX EA/OEA Final Chapter
- Page 83 and 84: COMPTUEX/JTFEX EA/OEA Final Chapter
- Page 85 and 86: COMPTUEX/JTFEX EA/OEA Final Chapter
- Page 87 and 88: COMPTUEX/JTFEX EA/OEA Final Chapter
- Page 89 and 90: COMPTUEX/JTFEX EA/OEA Final Chapter
- Page 91 and 92: COMPTUEX/JTFEX EA/OEA Final Chapter
- Page 93 and 94: COMPTUEX/JTFEX EA/OEA Final Chapter
- Page 95 and 96: COMPTUEX/JTFEX EA/OEA Final Chapter
- Page 97 and 98: COMPTUEX/JTFEX EA/OEA Final Chapter
- Page 99 and 100: COMPTUEX/JTFEX EA/OEA Final Chapter
- Page 101 and 102: COMPTUEX/JTFEX EA/OEA Final Chapter
- Page 103 and 104: COMPTUEX/JTFEX EA/OEA Final Chapter
- Page 105 and 106: COMPTUEX/JTFEX EA/OEA Final Chapter
- Page 107 and 108: COMPTUEX/JTFEX EA/OEA Final Chapter
- Page 109 and 110: COMPTUEX/JTFEX EA/OEA Final Chapter
- Page 111 and 112: COMPTUEX/JTFEX EA/OEA Final Chapter
- Page 113 and 114: COMPTUEX/JTFEX EA/OEA Final Chapter
- Page 115: COMPTUEX/JTFEX EA/OEA Final Chapter
- Page 119 and 120: COMPTUEX/JTFEX EA/OEA Final Chapter
- Page 121 and 122: COMPTUEX/JTFEX EA/OEA Final Chapter
- Page 123 and 124: COMPTUEX/JTFEX EA/OEA Final Chapter
- Page 125 and 126: COMPTUEX/JTFEX EA/OEA Final Chapter
- Page 127 and 128: COMPTUEX/JTFEX EA/OEA Final Chapter
- Page 129 and 130: COMPTUEX/JTFEX EA/OEA Final Chapter
- Page 131 and 132: COMPTUEX/JTFEX EA/OEA Final Chapter
- Page 133 and 134: COMPTUEX/JTFEX EA/OEA Final Chapter
- Page 135 and 136: COMPTUEX/JTFEX EA/OEA Final Chapter
- Page 137 and 138: COMPTUEX/JTFEX EA/OEA Final Chapter
- Page 139 and 140: COMPTUEX/JTFEX EA/OEA Final Chapter
- Page 141 and 142: COMPTUEX/JTFEX EA/OEA Final Chapter
- Page 143 and 144: COMPTUEX/JTFEX EA/OEA Final Chapter
- Page 145 and 146: COMPTUEX/JTFEX EA/OEA Final Chapter
- Page 147 and 148: COMPTUEX/JTFEX EA/OEA Final Chapter
- Page 149 and 150: COMPTUEX/JTFEX EA/OEA Final Chapter
- Page 151 and 152: COMPTUEX/JTFEX EA/OEA Final Chapter
- Page 153 and 154: COMPTUEX/JTFEX EA/OEA Final Chapter
- Page 155 and 156: COMPTUEX/JTFEX EA/OEA Final Chapter
- Page 157 and 158: COMPTUEX/JTFEX EA/OEA Final Chapter
- Page 159 and 160: COMPTUEX/JTFEX EA/OEA Final Chapter
- Page 161 and 162: COMPTUEX/JTFEX EA/OEA Final Chapter
- Page 163 and 164: COMPTUEX/JTFEX EA/OEA Final Chapter
- Page 165 and 166: COMPTUEX/JTFEX EA/OEA Final Chapter
COMPTUEX/JTFEX EA/OEA Final Chapter 3toothed dolphins in the SOCAL OPAREA; therefore seasonal occurrence can not be determined (Ferreroet al., 1994).Aco<strong>us</strong>tics—The vocal repertoire of the rough-toothed dolphin includes broad-b<strong>and</strong> clicks, barks, <strong>and</strong>whistles (Yu et al. 2003). Echolocation clicks of rough-toothed dolphins are in the frequency range of 0.1to 200 kHz, with a peak of about 25 kHz (Miyazaki <strong>and</strong> Perrin, 1994; Yu et al., 2003). Whistles show awide frequency range: 0.3 to >24 kHz (Yu et al., 2003). There is no published information on hearingability of this species.Short-beaked Common Dolphin (Delphin<strong>us</strong> delphis)Stat<strong>us</strong>—This is the most abundant cetacean off California (Dohl et al., 1981; Forney et al., 1995; Carrettaet al., 2005). The single current management unit for the short-beaked common dolphin in this area is aCalifornia/Oregon/Washington Stock, with a minimum population estimate of 449,846 (CV = 0.25)individuals (Carretta et al. 2004). The abundance of common dolphins varies seasonally but may beincreasing in California with a northward shift in the population (Heyning <strong>and</strong> Perrin, 1994; Barlow et al.,1997; Forney, 1997). The short-beaked common dolphin is not listed as endangered under the ESA or asdepleted or strategic under the MMPA.Distribution—Along the U.S. west coast, the short-beaked common dolphins’ distribution overlaps withthat of the long-beaked common dolphin. The short-beaked common dolphin is distributed between thecoast <strong>and</strong> at least 556 km from shore (Carretta et al., 2005). Short-beaked common dolphin abundanceoff California has increased dramatically since the late 1970s, along with a concomitant decrease inabundance in the ETP, suggesting a large-scale shift in the distribution of this species in the eastern NorthPacific (Forney et al., 1995; Forney <strong>and</strong> Barlow, 1998). The northward extent of short-beaked commondolphin distribution appears to vary inter-annually <strong>and</strong> with changing oceanographic conditions (Forney<strong>and</strong> Barlow 1998). Short-beaked common dolphins are found in the SOCAL OPAREA throughout theyear (Forney <strong>and</strong> Barlow, 1998).Stomach contents of Delphin<strong>us</strong> from California waters revealed 19 species of fish <strong>and</strong> 2 species ofcephalopods; Delphin<strong>us</strong> feeds primarily on organisms in the vertically migrating DSL (Evans, 1994).Diel fluctuations in vocal activity of this species (more vocal activity during late evening <strong>and</strong> earlymorning) appear to be linked to feeding on the DSL as it rises during the same time (Goold, 2000). Atagged individual tracked off San Diego conducted dives deeper than 200 m, but with most in the range of9 to 50 m (Evans, 1971, 1994).Aco<strong>us</strong>tics—Recorded Delphin<strong>us</strong> vocalizations include whistles, chirps, barks, <strong>and</strong> clicks (Ketten, 1998).Clicks <strong>and</strong> whistles have dominant frequency ranges of 23 to 67 kHz <strong>and</strong> 0.5 to 18 kHz, respectively(Ketten, 1998). Maximum source levels were approximately 180 dB 1 Pa-m (Fish <strong>and</strong> Turl, 1976).Oswald et al. (2003) found that short-beaked common dolphins in the ETP have whistles with a meanfrequency range of 6.3 kHz, mean maximum frequency of 13.6 kHz, <strong>and</strong> mean duration of 0.8 sec. Popov<strong>and</strong> Klishin (1998) recorded auditory brainstem responses from a common dolphin. The audiogram wasU-shaped with a steeper high-frequency branch. The audiogram b<strong>and</strong>width was up to 128 kHz at a levelof 100 dB above the minimum threshold. The minimum thresholds were observed at frequencies of 60 to70 kHz.Short-finned Pilot Whale (Globicephala macrorhynch<strong>us</strong>)Stat<strong>us</strong>—The short-finned pilot whale is not listed under the ESA. However, theCalifornia/Oregon/Washington Stock is considered strategic under the MMPA beca<strong>us</strong>e the averagehuman-ca<strong>us</strong>ed mortality may not be s<strong>us</strong>tainable (Barlow et al., 1997). A recent calculation of the3-53 February 2007