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Savory - Arachnida 1977

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184 Ill. PROLES ARACHNES<br />

24. THE ORDER OPILIONES 185<br />

FIG. 68. Phalangiurn, ventral aspect. After Roewer.<br />

FIG. 67. Legs ofOpiliones. After Kaestner. (i) Tarsus of Opilio parietinus; (ii) whole !eg<br />

of Nemasloma bimaculatum.<br />

The lower surface of the prosoma is formed from the labium, sternum<br />

and the coxae of the legs (Figs 68 and 69). The labium, as mentioned<br />

above, may be regarded as one of the mouth parts. It is probably homologous<br />

with the corresponding labium of Araneae and represents the<br />

sternite of one of the original somites. Behind it is the sternum proper, a<br />

single plate. In Laniatores it is long and narrow, reaching as far back as<br />

the fourth coxae. In Palpatores it is always short; sometimes it is continuous<br />

with the labial portion and sometimes with the genital plates<br />

behind it, but it is often largely out of sight, hidden below the coxae.<br />

The coxae are sometimes all immovable and sometimes all movable in<br />

Palpatores; in Laniatores the three posterior pairs are immovable, the<br />

fourth pair coalescing with the opisthosoma, while the first pair can be<br />

rotated, moving their maxillary lobes towards or away from the<br />

mandibles.<br />

The opisthosoma is usually so closely united to the prosoma that the<br />

distinction between the two parts is not obvious. Its precise structure is<br />

not easily elucidated without the comparison of many forms, since segments<br />

are often fused together or are missing, and also because the<br />

tergites and. sternites of the same segment are not always placed opposite<br />

each ~ther m the lateral aspect of the adult. The shape of the opisthosoma<br />

IS usually oval or globular, but it is flattened in Trogulidae. It is<br />

covered with a fairly hard exoskeleton often decorated with spines. In<br />

European species these are small and usually amount to no more than<br />

rows of points indicating segments, but in many tropical forms as in the<br />

Gonyleptidae, there are more elaborate developments. '<br />

In most Palpatores, nine tergites and nine sternites can be recognized,<br />

followed by the operculum anale, a small circular plate pierced by a<br />

transverse or round anus. This operculum is regarded as the tergite<br />

only of the tenth somite. Hence in these two sub-families there are ten<br />

FIG. 69. Hornalenotus, ventral aspect, expanded.

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