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Savory - Arachnida 1977

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38 II. DE ARACHNIDIS<br />

4. PHYSIOLOGY: II'\TER:KAL ORGANS 39<br />

m IV V<br />

i<br />

~---- -------~------------~------------- --------- _,--,___<br />

ii<br />

FIG. 11. i\ervous system of a spider.<br />

This is regarded as the more primitive method. In some orders it<br />

has been replaced by adapting a limb for conveying the sperm: the<br />

pedipalpi in Araneae, the third leg in Ricinulei. Lastly, there may be a<br />

real intromittent organ and copulation in the true sense may occur, as<br />

in the Opiliones and some Acari.<br />

Three distinct methods of excretion are found among <strong>Arachnida</strong>,<br />

the most fully developed of which is the system of coxal glands (Figs 13<br />

and 14). In their typical form, many modifications of which are,<br />

however, found, they consist of a large excretory saccule, lined with<br />

---------~- -------~--------~iii<br />

--------~-<br />

FIG. 13. Coxal glands. (i 1 Scorpiones; >-\mblyjlygi; (iiii >\mbly;Jygi !Charontini!;<br />

Uropygi; (v', Pal;'igradi.<br />

FIG. 12. Diagram of female reproductive organs of a spider. A, ovary; B, oviduct; C,<br />

spermatheca; D, orifice for entry of sperm; E, vagina.<br />

cubical or flattened epithelium, lying outside the cndosternitc opposite<br />

the coxae of the first legs. This discharges its products into a eo m oluted<br />

tube, the labyrinth, the coils of which occupy the space from the first to<br />

the fourth coxa, or beyond. At the distal end of the labyrinth there is<br />

sometimes a S\n·lling, rt>garded as a "bladder". and from this there runs<br />

forward a straight tube, the internal limb of the labyrinth, which lies

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