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Savory - Arachnida 1977

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266 IV. DE ARACHNOLOGIA<br />

little was known about them, he produced a logical system of the order,<br />

and in addition to many papers summarized his system in a great<br />

monograph "The Arachnid Order Chelonethida" in 1931.<br />

For many years arachnologists the world over looked to Alexander<br />

Petrunkevitch (1875-1964) as the supreme authority on all questions of<br />

arachnid evolution and classification. Born in Russia, he travelled to<br />

America in 1903 and in 1910 joined the University of Yale. His attempts<br />

to compile a satisfactory classification of spiders, his investigation of<br />

extinct fossil species and his later work on spiders in amber were the<br />

chief items in a long life characterized by an incredible daily schedule<br />

of hours in the laboratory, remarkable technical skill and an astonishing<br />

command of about ten European languages. The influence of his<br />

contributions to arachnology will survive for many years to come.<br />

Africa, with its great fauna of conspicuous scorpions and Solifugae,<br />

has always offered boundless opportunities to arachnologists. After a<br />

few casual references to spiders among various descriptions of insects<br />

there appeared in 1825 the well known "Description de l'Egypte" by<br />

]. S. Savigny and V. ]. Audouin. Nine plates, engraved by Savigny,<br />

accompanied descriptions of 29 genera of spiders, together with some<br />

account of other orders. Indeed, the history of both scorpions and<br />

Solifugae makes constant references to the African species; and from<br />

Africa too came in 1838 the recognition of the first of the Ricinulei, the<br />

species which at that time was known as Cryptostemma westermannii.<br />

Most of the study of the African <strong>Arachnida</strong> has come from the southern<br />

half of the continent. The pioneer work must be credited to W. P.<br />

Purcell (1866-1919),]. Hewitt (1880-) and E. Warren (1871-1935),<br />

all of whom were born in Britain and spent a large part of their lives in<br />

Africa. Employed in the service of different museums, they were the<br />

pioneers of arachnology in Africa.<br />

Here it is impossible to omit mention of a living arachnologist, Dr<br />

R. F. Lawrence. In 1922 he accepted a post in the South African<br />

Museum at Cape Town, where Purcell's large collections were preserved.<br />

In the following years he undertook extensive expeditions to<br />

various parts of the continent, returning, whether from mountain,<br />

forest or desert, with a full harvest of specimens. In 1935 he succeeded<br />

Warren as Director of the Natal ~1useum at Pietermaritzburg, and in<br />

1953 published his impressive volume on "The Cryptic Fauna of<br />

Forests". Lawrence's work has covered almost all the orders of arachnids,<br />

and he must be recognized as having securely laid the foundations<br />

of African arachnology.<br />

In the Near, Middle and Far East, arachnology has made steady if<br />

31. HISTORICAL ARACHNOLOGY 267<br />

unspectacular progress during the present century. In these regions,<br />

as indeed in most parts of the world, its growth has followed the same<br />

course. At first a few curious examples of one order or another have been<br />

rather vaguely mentioned in a book of travels, and from these there<br />

grew some interest in the local fauna. More serious collections were<br />

made, often by explorers or professional collectors, and were brought<br />

home to specialists in Britain, France or Germany. In this way a very<br />

large number of foreign <strong>Arachnida</strong> were described by zoologists such as<br />

R. I. Pocock (1863-1947), F. H. Graveley (1885- ), vV. R. Sherriffs<br />

(1881- ) and H. R. Hogg (1851-1923).<br />

This period overlaps the later stage of development, when the inhabitants<br />

of a country, provided with museums and universities of their<br />

own, undertook these duties themselves. By progress of this kind there<br />

has grown an increasing concern with <strong>Arachnida</strong> in India, Palestine,<br />

Malaysia, Japan and elsewhere.<br />

In the Antipodes the study of the spiders of New Zealand began with<br />

a collection made in 1827 during the voyage of the Astolabe. This was<br />

followed during the next 50 years by a small number of other collections,<br />

one of which included the species locally known as the Katipo,<br />

notorious for its poisonous bite. This was a species of Latrodectus, a<br />

relation of the famous black widow.<br />

The true founder of ~ew Zealand arachnology was A. T. Urquhart<br />

(1839-1916), a keen naturalist who made good use of his travels in the<br />

country. His many papers were published between 1882 and 1897, and<br />

together they gave a sound start to the subject. Some years passed<br />

before another took up the study. This was Comte R. de Dalmas ( 1862-<br />

1930). At first he brought his many specimens to Simon for repo~t, but<br />

later he examined and described them himself.<br />

The thorough and systematic description of the <strong>Arachnida</strong> of New<br />

Zealand, now in the hands of R. R. and L. M. Forster, is of great<br />

significance, since it has shown how the taxonomy of <strong>Arachnida</strong> has<br />

been handicapped by the fact that all systems have been based on<br />

specimens from the northern hemisphere, to the exclusion of the often<br />

very different fauna of the other half of the world.<br />

The first <strong>Arachnida</strong> from Australia were briefly mentioned by]. C.<br />

Fabricius in his "Entomologica Systematica" of 1792-96. Subsequent<br />

collections were described by L. Koch and Graf von Keyserling in a<br />

great work of nearly 1,500 pages, "Die Arachnid en Australiens". In<br />

the present century many Australian species were examined by H. R.<br />

Hogg. The first resident in Australia to continue this work was W. ].<br />

Rainbow (1856-1919). Born in Yorkshire, he had lived in New Zealand<br />

and Australia from 1873, and 20 years later became entomologist to the

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