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Savory - Arachnida 1977

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!,<br />

240 Ill. PROLES ARACHNES<br />

29. THE ORDER SOLIFUGAE 241<br />

diameter carries setae 3 cm long. It is said that if the tip of but a single<br />

seta be touched even with a hair, the animal responds instantaneously.<br />

The opisthosoma of Solifugae is always a regular oval, the posterior<br />

margin being a smooth curve with no trace of a post-anal structure. It is<br />

always conspicuously and completely segmented, and the somites<br />

carry both tergites and sternites. The number of somites appears to he<br />

ten, but 11 are present in the embryo, and this number is really retained<br />

in the adult, the first somite being much reduced. Its sternite<br />

is a small triangular plate between the fourth coxae, and its tergite is<br />

similar in shape and lies hidden between the prosoma and the opisthosoma.<br />

The genital orifice on the second sternite is a slit-like aperture<br />

peculiar among <strong>Arachnida</strong> in being guarded by movable lips, so that<br />

it can open and close. The third, fourth and fifth sternites show just<br />

behind their posterior edges the orifices of the opisthosomatic tracheae.<br />

There is a pair of these behind the third and the fourth sternites and a<br />

single, median one behind the fifth.<br />

DISTRIBUTION<br />

Solifugae are confined mainly to the tropical and sub-tropical regions,<br />

and in Europe occur only in the south-east of Spain (Fig. 105). The<br />

distribution of the separate families may be summarized thus:<br />

FAMILY<br />

DISTRIBUTION<br />

Fw. 104. Solifugae; ventral aspect.<br />

Galeodes arabs.<br />

differ from that of all others. The first pair of are much reduced<br />

and bear only three racket-organs each. Since the young of other<br />

families possess three such organs, this suggests that the Hexisopodidae<br />

exhibit neoteny. They and the family Rhagodidae have legs that are<br />

better adapted to digging than the legs in other families. The legs in all<br />

the Solifugae bear spines or setae of every degree of stoutness from short<br />

sharp spikes to long hair-like setae. There is not much evidence of<br />

thinking that any of these have any other function than that of organs<br />

of touch, as in all the orders of the <strong>Arachnida</strong>, but in Solifugae they<br />

seem to be more plentiful and more sensitive than in their allies. No<br />

other arachnid is so ''hairy" that to a casual glance it looks like a ball of<br />

fluff, and no other can show such proportions as those found, for examplc,<br />

in the fourth leg of the male Solpuga monteiroi, where a leg 2 mm in<br />

Melanoblossiidae<br />

Eremobatidae<br />

Karschiidae<br />

Rhagodidae<br />

Hexisopodidae<br />

Gylippidae<br />

Solpugidae<br />

Ammotrechidae<br />

Galeodidae<br />

Ceromidae<br />

Amacataidae<br />

Dacsiidae<br />

South-East<br />

?\ortb and Central America<br />

Asia; l'\ car East; north -west Africa<br />

north-east Africa; Near East; south-west Africa<br />

South Africa<br />

East Asia<br />

Africa<br />

South and Central America; southern North America<br />

Asia; North Africa<br />

South Africa<br />

western South America<br />

Near East; Africa; 1\:feditcrrancan zone<br />

The above table may be expanded thus. Solifugae occur over almost<br />

the whole of Africa. In Asia they are to be found in Arabia, Persia,<br />

Turkestan and India. They also occur in Celebes and Indo-China. The<br />

:r\ ear East species extend as far west as the coast of Palestine and as far<br />

north as the Caspian Sea.<br />

American Solifugae fully studied by M. H. Muma belong to the<br />

families Eremobatidae and Ammotrechidae. They are now known from

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