Vol. 16—1962 - NorthEastern Weed Science Society

Vol. 16—1962 - NorthEastern Weed Science Society Vol. 16—1962 - NorthEastern Weed Science Society

08.06.2015 Views

16 SOMECHARACTERISTICS OF QUACKQR4SS ArID THEIRRELATIONTO CONTROLY '.. . '.' .',,'> ',".' ,I " ,'.' ,', K. P. ElUchholtz y' ·,_,.i" .- QuaeJcgrass is a widely distributed end persietent'weedili the northern states arid Canada. .It has proved to be adapted to 'Cobt,. temperate n!gions and lsfevoi'ed .by' hUmid or subJ.!tIum1dcUinstell." 1he;weed is IOOsttroubl8some in areas thatdare subjected tcqlerlodic 1 but'no,t; ,cxmtinuous tillage. It Competes stronglY with forage cropS',; gn-ins, com··1tt!d'other row crops', f'J:'lrl,ts, and: vegetables. !:t is objectionable in forestry p1:antations, laws, 'llndindustria~ sites. In ordert,(, develop effecti.~Controlprogr,.ror qusckgra ss it, i8 necessary ,to' consider the fectors that account,tb-r1ts adaptation in,.the' northe~ states and the reaso,ns for its pers;l.stene;:e•. ' Quackgrass spre8~') by both rb1'ZOIlIeS, and seeds. It iSl'robehlethat ~ad by seeds is J1I!)rj:_ w;idespreadthanis generally rElaJ;tzed. Because or':aimi.larity in shape,t,!'1~ seed is more cOll!lOOn in oats than itl seed of other'icereals. In 1957 a stUdy in Wisconsin showed that 35%of 799 oat samples handled by the State Seed Testing Laboratory in that year contained qusckgrass seeds. Seeder box surveys of oats being seeded have consistently shown that from 50~ to 60%of the grain being used as seed in Wisconsin is infested with qusckgrass seed. Viability of quackgrass seed in seed grain was frequently as high as that of the oats. In addition to the spread in seed grain the mature seeds of quackgrass are frequently harvested with hay and ultimately find their way back to the field in manure or bedding. Seeds also mature in pastures where they shatter readily and infest the soil. The prevalence and vJ.abil1ty of qusckgrass seeds often allows reinfestetion aftsr all established plants hava been eradicated. The ability of quackgrass to spread by rhizomes is widely recognized. These spreading underground stems may extend as 1IIUChas 3 or 4 feet in a single year. Tillage often extends the area of infestation IllUchfarther than this by dragging fragments of the rhizomes beyond the area of initial infestation. Rhizome production is prolific in a productive soil. Meyer (9), Schirman and Buchholtz (14) and Johnson (4) all found as much as 6000 lblA of dry rhizomes in heavily infested areas. This mass of rhizomes was contained in the plow slice and in an undisturbed sod was concentrated in the upper 4 inches of so11. The life of a particular rhizome is not as long as would be expected. Sagar (13) mentioned that the rhizomes may live for 3 years in England, but seldom longer. Johnson and Buchholtz (6) found no evidence that the rhizomes lived more than 2 years under IOOrerigorous conditione in Wisconsin and a large proportion of them lived no longer than 1 year. The fact that the rhizomes are relatively short-lived is a vulnerable point in the life cycle of the weed. If new rhizome growth could be limited or eliminated for 1 year, the persistence of the weed would be sharply reduced. 11Published with the approval of the Director of the Wisconsin Agricultural Experiment Station as a collaborator under Horth Central Regional Coop- ___ ..1.,, '1"\__ .I. __ ...L '-1_ "In

The vigorous regrowth of quackgrass is related to the large number of dormant buds that are present on the r!;lizomes in an in;f'ested area. Weight of dry quackgrass rhizomes averages about 0.4 gm per foot.' If an area contains $000 Ib of the dry rhizomes per acre, this is equal to about 130 feet of rhizome per sq ft. The nodes on rhieomes average no more than 1 inch apart. Therefore, it is safe to assume that in excess of 1500 buds exist on the rhizomes in each sq !t of infested soil. Quackgrass standl1 seldom exceed '::tSOshoots per sq ft. From this it is apparent that 90 percent or more of the bilds on the rhizomes retn8in inactive until circumstances are favorable for their development. This reservoir of dormant bUds is a major hazard, for wheneyer. a stan~ of shoots is destroyed, some of the dormant buds Will be activated' and will very soon res13ta - blish the plant. . . The rhizomes Of qua.ckgrass contain a number of carbohydra.tes of which trit1c1n is probably present in greatest concentration. Total available carbj'hydrates, vary from about 30 to $0%of the dry rhizom~tweight according to Pinckney (12), Schirman and Buchholtz (1.4) and LeBaron and F~rtig (7). If one aSflWnesan average concentration of availa9le carbohydrates of ,40~ and a total dry ma~ter content of $000 lb/A, it appears that a reserve of 2000 lb/A of readily av~lable carbohydrates is stored in the rhizomes. Such,li!large reserve, along with the abundant supply of inactive bu4s, is ample reason why the growth of quackgr8J.ss is vigorous and w!:lythe regrowth occurs with such persistence. . The trends in total carbohydrate reserves during the year in quackgrass do not show the fluctuations that are found in many hel;'baceous plants. The work of Pinckney (12) and Schirman and Buchholtz (14) show;s that only minor var;Lations in reserve level occur during the year. LeBaron and, FeX'tig (7) found a substantial reduction in fructose content of the rhizomes djl.;r'ingthe winter, but their data are not fully comparable to the others cited, for it is based only on ,the content of fructose found in th~. tissue. Since there appeare to be no marked depression in total carbohydrate level during the seaeon, there is no most favorable time at which control measures directed at reducing the carbohydrate content may be initiated. Stoa, et aL, (15) was one of the first to comment,that the carbohydrate reserves of quackgrasa could be reduced by continual and consistent defoliat10n. Dexter (2) showed that organic reserves were reduced more readily after fertilization with nitrogen~ Defoliation by some tillage 9peration is no doubt effective, but it is inconvenient and requires the majqr portion of a season to produce the desired results. Alternative methods involving use of herbicides now'appe'ar l!I)re promising. Buchholtz (1), M~ggitt (8) and Fertig (3) have all reported on the superior control of quackgrass obtained withatrazine and other triazine herbicides.According to Schirman and Buchholtz (14) and LeBaron and Fertig (7) control results from the drastic reduction in carbohydrate reserves in the rhizomes following the treatment. !tappears that respiratory activity is maintained at a high l,-r'lll after trea'\;ment even t.hough photosynthesis is interrupted. The result is ~_.faster rate of depletion than can be accomplished by consistent defoliation or,m . other known method. D~pletion appears to proceed more rapidly when the t~, growth of the quackgrass is allow-edto remain intact for a month or more following treatment than.when it is removed. This is reasorulble for the presence:of the top growth increases the total respiratory activ,Lty of the plant. j, ' 17

16<br />

SOMECHARACTERISTICS OF QUACKQR4SS ArID THEIRRELATIONTO CONTROLY<br />

'.. . '.' .',,'> ',".' ,I " ,'.' ,',<br />

K. P. ElUchholtz y' ·,_,.i"<br />

.-<br />

QuaeJcgrass is a widely distributed end persietent'weedili the northern<br />

states arid Canada. .It has proved to be adapted to 'Cobt,. temperate n!gions<br />

and lsfevoi'ed .by' hUmid or subJ.!tIum1dcUinstell." 1he;weed is IOOsttroubl8some<br />

in areas thatdare subjected tcqlerlodic 1 but'no,t; ,cxmtinuous tillage. It Competes<br />

stronglY with forage cropS',; gn-ins, com··1tt!d'other row crops', f'J:'lrl,ts,<br />

and: vegetables. !:t is objectionable in forestry p1:antations, laws, 'llndindustria~<br />

sites.<br />

In ordert,(, develop effecti.~Controlprogr,.ror qusckgra ss it, i8<br />

necessary ,to' consider the fectors that account,tb-r1ts adaptation in,.the'<br />

northe~ states and the reaso,ns for its pers;l.stene;:e•. ' Quackgrass spre8~')<br />

by both rb1'ZOIlIeS, and seeds. It iSl'robehlethat ~ad by seeds is J1I!)rj:_<br />

w;idespreadthanis generally rElaJ;tzed. Because or':aimi.larity in shape,t,!'1~<br />

seed is more cOll!lOOn in oats than itl seed of other'icereals. In 1957 a stUdy<br />

in Wisconsin showed that 35%of 799 oat samples handled by the State Seed<br />

Testing Laboratory in that year contained qusckgrass seeds. Seeder box<br />

surveys of oats being seeded have consistently shown that from 50~ to 60%of<br />

the grain being used as seed in Wisconsin is infested with qusckgrass seed.<br />

Viability of quackgrass seed in seed grain was frequently as high as that<br />

of the oats. In addition to the spread in seed grain the mature seeds of<br />

quackgrass are frequently harvested with hay and ultimately find their way<br />

back to the field in manure or bedding. Seeds also mature in pastures where<br />

they shatter readily and infest the soil. The prevalence and vJ.abil1ty of<br />

qusckgrass seeds often allows reinfestetion aftsr all established plants<br />

hava been eradicated.<br />

The ability of quackgrass to spread by rhizomes is widely recognized.<br />

These spreading underground stems may extend as 1IIUChas 3 or 4 feet in a single<br />

year. Tillage often extends the area of infestation IllUchfarther than this<br />

by dragging fragments of the rhizomes beyond the area of initial infestation.<br />

Rhizome production is prolific in a productive soil. Meyer (9), Schirman<br />

and Buchholtz (14) and Johnson (4) all found as much as 6000 lblA of dry<br />

rhizomes in heavily infested areas. This mass of rhizomes was contained in<br />

the plow slice and in an undisturbed sod was concentrated in the upper 4<br />

inches of so11.<br />

The life of a particular rhizome is not as long as would be expected.<br />

Sagar (13) mentioned that the rhizomes may live for 3 years in England, but<br />

seldom longer. Johnson and Buchholtz (6) found no evidence that the rhizomes<br />

lived more than 2 years under IOOrerigorous conditione in Wisconsin and a<br />

large proportion of them lived no longer than 1 year. The fact that the rhizomes<br />

are relatively short-lived is a vulnerable point in the life cycle of<br />

the weed. If new rhizome growth could be limited or eliminated for 1 year,<br />

the persistence of the weed would be sharply reduced.<br />

11Published with the approval of the Director of the Wisconsin Agricultural<br />

Experiment Station as a collaborator under Horth Central Regional Coop-<br />

___ ..1.,, '1"\__ .I. __ ...L '-1_ "In

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