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Paleozoic Voltziales

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Fossil History of Plants<br />

Lecture 12<br />

Rise of the conifers - <strong>Paleozoic</strong><br />

<strong>Voltziales</strong>


Five major groups of vascular plants


Not Big<br />

Like these<br />

Mesozoic Conifers<br />

Meanwhile on the dryer slopes


Walchian conifers (<strong>Voltziales</strong>)<br />

(2 – 3 M tall)<br />

small<br />

Originally named Lebachia –Now Emporia and others


<strong>Voltziales</strong><br />

Walchian conifers<br />

U. Penn. & L. Permian<br />

conifers


Emporia leafy shoot<br />

and<br />

woody stem


Emporia seed cone and seeds<br />

with embryos


Voltzialean Conifer Phylogeny


Walchian conifer leafy shoots<br />

Hanskerpia


Walchian conifer leafy shoot anatomy<br />

Hanskerpia


Walchian conifer leaf structure and anatomy<br />

Hanskerpia


Walchian conifer (Barthelia)


Ideas of conifer cone evolution from Rudolf Florin 1930’s-1964


Conifer cone evolution and phylogeny<br />

R. Florin – 1930s-1960s<br />

Missippian & Pennsylvanian<br />

Cordaitales<br />

Pennsylvanian to Permian<br />

Conifers (<strong>Voltziales</strong>)<br />

Jurassic/Cretaceous to Recent<br />

Conifers (modern families)<br />

Gifford & Foster, 1989


a<br />

Pennsylvanian<br />

Permian<br />

Triassic<br />

Jurassic<br />

*<br />

Cretaceous<br />

Eocene<br />

Fossils,<br />

Stratigraphy<br />

&<br />

Conifer<br />

Phylogeny<br />

*<br />

*


Pattern of Modern Conifer Phylogeny<br />

Living plants Fossil Record<br />

Rai et al., 2008<br />

Triassic<br />

Jurassic<br />

Triassic?<br />

Cretaceous

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