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A Review of the Genus Eunice - Smithsonian Institution Libraries

A Review of the Genus Eunice - Smithsonian Institution Libraries

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252 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY<br />

paired, yellow, distally truncate or gently hammer-headed<br />

(Figure 83b,e); cross-section round. Separation <strong>of</strong> cores and<br />

sheaths indistinct in both aciculae and subacicular hooks.<br />

Subacicular hooks (Figure 83c) yellow, tridentate with teeth in<br />

a crest. Hooks first present from setiger 21. Hooks with<br />

well-developed, distinct fangs.<br />

UNKNOWN MORPHOLOGICAL FEATURES.—Relationship between<br />

Mx III and left Mx IV; pygidium and anal cirri.<br />

EXPECTED STATES OF UNKNOWN MORPHOLOGICAL FEA-<br />

TURES.—None.<br />

CHARACTERS USED IN PREPARATION OF KEY NOT<br />

SCORED.—Inappropriate Characters: 34, 56. Unknown<br />

Characters: 1, 2,4,6, 32,42, 47, 50, 57-60, 63, 81, 82.<br />

ASSUMED STATES FOR PURPOSE OF PREPARING KEY.—<br />

None.<br />

REMARKS.—<strong>Eunice</strong> papeetensis was described as a subspecies<br />

<strong>of</strong> E. oliga; it differs at <strong>the</strong> species level from that species<br />

in that it has branchiae limited to a short anterior region; £.<br />

oliga has branchiae continued to near <strong>the</strong> posterior end.<br />

<strong>Eunice</strong> papeetensis is listed with similar species in Tables 41<br />

and 44. It has maximally three branchial filaments, whereas all<br />

o<strong>the</strong>r species in Table 44 have at least eight filaments where <strong>the</strong><br />

branchiae are best developed.<br />

146. <strong>Eunice</strong> parasegregata Hartmann-Schroder, 1965<br />

FIGURE 83g-n; TABLES 19,21-23<br />

<strong>Eunice</strong> parasegregata Hartmann-Schroder, 1965b:168-172, figs. 140-144.<br />

MATERIAL EXAMINED.—Holotype, ZMH P-14273, <strong>of</strong>f<br />

Punta Lavapie, Chile, 37°08.7'S, 73°38.6'W, 58 m, fine sand<br />

with rocks and dead algae, dredge and grab, 10 Mar 1960, coll.<br />

Exp. Mar. Chile I.<br />

COMMENTS ON MATERIAL EXAMINED.—The holotype is<br />

currently in three pieces: an anterior end <strong>of</strong> 33 setiger, a median<br />

piece <strong>of</strong> 10 branchiated setigers, and a mid-posterior abranchiate<br />

piece <strong>of</strong> 22 setiger. A number <strong>of</strong> segments are missing<br />

between both <strong>the</strong> anterior end and <strong>the</strong> short median fragment<br />

and between <strong>the</strong> latter and <strong>the</strong> medioposterior fragment, but<br />

exactly how many segments are missing cannot currently be<br />

estimated.<br />

DESCRIPTION.—Anterior end with 33 setigers; length 22<br />

mm; maximal width 3 mm; length through setiger 10, 8 mm.<br />

Prostomium (Figure 83g) distinctly shorter and narrower<br />

than peristomium, as deep as x li <strong>of</strong> <strong>the</strong> peristomium. Prostomial<br />

lobes frontally rounded, dorsally inflated; median sulcus<br />

shallow. Eyes under peristomial fold, posterior to bases <strong>of</strong> A-I,<br />

brown. Antennae in a horseshoe, evenly spaced, similar in<br />

thickness. Ceratophores ring-shaped in all antennae, without<br />

articulations. Ceratostyles slender and tapering; A-I with 3-4<br />

drop-shaped articulations; A-I I and A-I 11 with 5 to 6 irregular,<br />

cylindrical articulations; all articulations on distal half <strong>of</strong><br />

ceratostyles. A-I to posterior peristomial ring; A-II to setiger 5;<br />

A-I 11 to setiger 6. Peristomium cylindrical. Separation between<br />

distinct on all sides; anterior ring twice as long as posterior<br />

ring. Peristomial cirri to middle <strong>of</strong> prostomium, slender and<br />

digitiform, without articulations.<br />

Maxillary formula 1+1, 14+16, 13+0, 10+16, and 1+1. Mx<br />

III long, located behind left Mx II. Mx VI missing.<br />

Branchiae (Figure 83n) present, pectinate, distinctly longer<br />

than notopodial cirri, not reduced in mid-body region, erect.<br />

Branchiae from setiger 3- to end <strong>of</strong> fragment, present on all<br />

setigers <strong>of</strong> median fragment, missing on medioposterior<br />

fragment. Branchiae terminating well before posterior end. All<br />

branchiae with 2 or more filaments; maximum 18 filaments at<br />

setiger 10; branchiae on median fragment with 2 to 3 filaments;<br />

those <strong>of</strong> last segments in anterior end with at least 7 filaments.<br />

All filaments slender, tapering, as long as notopodial cirri<br />

except in first branchial segment.<br />

Anterior ncuropodial acicular lobes distally obliquely truncate,<br />

becoming distinctly truncate in last segments present;<br />

aciculae emerging dorsal to midline. In medioposterior<br />

fragment acicular lobes reduced, equally supported by aciculae<br />

and subacicular hooks (Figure 83m). Anterior pre- and<br />

postsetal lobes low, transverse folds. Posterior pre- and<br />

postsetal lobes follow outline <strong>of</strong> acicular lobes closely. First 5<br />

ventral cirri thick, tapering, becoming distinctly basally<br />

inflated in branchiated setigers. Inflated bases nearly spherical;<br />

narrow tips digitiform. Postbranchial ventral cirri slender and<br />

nearly digitiform. All notopodial cirri basally slightly inflated,<br />

tapering to slender tips, without articulations.<br />

Limbate setae slender, marginally smooth. Pectinate setae<br />

(Figure 83i,l) tapering, flat One marginal tooth much longer<br />

and heavier than o<strong>the</strong>r teeth, -10 teeth present. Shafts <strong>of</strong><br />

compound falcigers (Figure 83hjc) distally inflated, marginally<br />

serrated. Appendages triangular, bidentate. Proximal teeth<br />

smaller than distal teeth, triangular, directed laterally. Distal<br />

teeth gently curved. Guards distally asymmetrically bluntly<br />

pointed; mucros absent Pseudocompound falcigers and compound<br />

spinigers absent. Aciculae yellow, distally tapering,<br />

gently curved; cross-section round. Separation between cores<br />

and sheaths indistinct in both aciculae and subacicular hooks.<br />

Subacicular hooks (Figure 83j) yellow, bidentate. Hooks<br />

present on median and medioposterior fragments, present in all<br />

setigers, always single (except for replacements). Hooks<br />

tapering. Proximal teeth larger than distal teeth, directed<br />

laterally. Distal teeth very short, nearly erect.<br />

UNKNOWN MORPHOLOGICAL FEATURES.—Parapodial and<br />

setal features <strong>of</strong> far posterior setigers; pygidium and anal cirri.<br />

EXPECTED STATES OF UNKNOWN MORPHOLOGICAL FEA-<br />

TURES.—None.<br />

CHARACTERS USED IN PREPARATION OF KEY NOT<br />

SCORED.—Inappropriate Characters: 56, 58, 59. Unknown<br />

Characters: 1, 2,4,6, 36, 38, 40, 80.<br />

ASSUMED STATES FOR PURPOSE OF PREPARING KEY.—38,2.<br />

REMARKS.—<strong>Eunice</strong> parasegregata may belong to ei<strong>the</strong>r<br />

group A-l or A-2 and has been listed in Tables 19 and 22 as<br />

well as in Tables 21 and 23. Because <strong>of</strong> <strong>the</strong> fragmented state <strong>of</strong>

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