A Review of the Genus Eunice - Smithsonian Institution Libraries
A Review of the Genus Eunice - Smithsonian Institution Libraries
A Review of the Genus Eunice - Smithsonian Institution Libraries
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188 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY<br />
SCORED.—Inappropriate Characters: 56, 58, 59. Unknown<br />
Characters: 1,2,4,6, 13, 14, 42,47, 50,63.<br />
ASSUMED STATES FOR PURPOSE OF PREPARING KEY.—<br />
None.<br />
REMARKS.—<strong>Eunice</strong> kobiensis was considered a synonym <strong>of</strong><br />
E. websteri (as E. longicirrata Webster) by Imajima and<br />
Hartman (1964:256); as already indicated by Fauchald (1969),<br />
<strong>the</strong> proposed synonymy cannot be correct. <strong>Eunice</strong> kobiensis is<br />
compared to similar species in Tables 19 and 20. Fauchald<br />
(1969) compared it to a number <strong>of</strong> similar species; at this time<br />
nothing can be added to that comparison and to discussions <strong>of</strong><br />
o<strong>the</strong>r species listed in Table 20.<br />
99. <strong>Eunice</strong> langi (Treadwell, 1943)<br />
FIGURE 61j-o; TABLES 24,26<br />
Leodice langi Treadwell, 1943:3, figs. 14-18.<br />
<strong>Eunice</strong> savignyi.—Hartman, 1956:283 [not <strong>Eunice</strong> savignyi Grube, 1878].<br />
MATERIAL EXAMINED.—Holotype, AMHN 3739, Cape<br />
Town, South Africa.<br />
COMMENTS ON MATERIAL EXAMINED.—The anterior end<br />
has been badly mangled and partly dissected, and all antennae<br />
are lost, so no illustrations were attempted <strong>of</strong> <strong>the</strong> anterior end.<br />
DESCRIPTION.—Holotype complete, <strong>of</strong> unknown sex, with<br />
71 setigers; total length 42 mm; maximal width 2 mm; length<br />
through setiger 10, 5.5 mm.<br />
Prostomial lobes frontally obliquely truncate, dorsally<br />
flattened; median sulcus shallow. Eyes not observed. Ceratophores<br />
ring-shaped in all antennae. Ceratostyles missing.<br />
Peristomial rings distinct dorsally and ventrally; anterior ring<br />
3 A <strong>of</strong> total peristomial length. Peristomial cirri to setiger 3, with<br />
6 distinct articulations.<br />
Maxillary formula 1+1,6+5,9+0,6+8, and presumably 1+1<br />
according to Treadwell (1943, fig.18).<br />
Branchiae (Figure 61k) present, pectinate, distinctly longer<br />
than notopodial cirri, not reduced in mid-body region, erect.<br />
Branchiae from setiger 3 to setiger 39. Maximum 8 filaments at<br />
or near setiger 15; most branchiae with 4 or 5 filaments. Stems<br />
stout, strongly tapered, shorter than notopodial cirri. Filaments<br />
digitiform, about as long as notopodial cirri.<br />
Anterior neuropodial acicular lobes distally truncate; posterior<br />
acicular lobes (Figure 61j) obliquely conical; aciculae<br />
emerging at midline. Pre- and postsetal lobes low, transverse<br />
folds. First 4 ventral cirri very long, digitiform, as long as<br />
notopodial cirri. Ventral cirri basally inflated from setiger 5 to<br />
setiger 36, with distinct, digitiform, obliquely attached tips.<br />
Inflated bases increasing evenly from setiger 5 to setiger 15,<br />
ovate, decreasing from setiger 30, missing posterior to setiger<br />
36. Posterior ventral cirri short, nearly tubercular. Anterior<br />
notopodial cirri slender, digitiform; posterior notopodial cirri<br />
long, tapering. Notopodial cirri without articulations.<br />
Shafts <strong>of</strong> pectinate setae (Figure 61m) slender, cylindrical.<br />
Blades flat, strongly flaring. One marginal tooth longer than<br />
o<strong>the</strong>r teeth, with -15 teeth. Shafts <strong>of</strong> compound falcigers<br />
(Figure 611) gently inflated, marginally smooth. Appendages<br />
long, tapering; heads very small, bidentate. Proximal teeth<br />
much smaller than distal teeth, very short, triangular, directed<br />
laterally. Distal teeth tapering, nearly erect. Guards asymmetrically<br />
sharply pointed; mucros absent. Pscudocompound falcigers<br />
and compound spinigers absent. Aciculae (Figure 61n)<br />
single, amber-colored, tapering, distinctly bent in median and<br />
posterior setigers; cross-sections round. Subacicular hooks<br />
(Figure 61o) amber colored, bidentate. Hooks first present from<br />
setiger 30, present in all setigers <strong>the</strong>reafter, always single<br />
(except for replacements). Hooks tapering; heads small,<br />
distinctly curved. Proximal and distal teeth similar in size.<br />
Proximal teeth directed laterally, blunt. Distal teeth truncate,<br />
directed laterally.<br />
UNKNOWN MORPHOLOGICAL FEATURES.—Prostomial features;<br />
relationship between Mx III and left Mx IV; pygidium<br />
and anal cirri.<br />
EXPECTED STATES OF SELECTED UNKNOWN FEATURES.—<br />
Mx III short; forming a distal arc with left Mx IV.<br />
CHARACTERS USED IN PREPARATION OF KEY NOT<br />
SCORED.—Inappropriate Characters: 56, 58, 59. Unknown<br />
Characters: 4, 6, 10-17, 19-24, 39-40,74, 78.<br />
ASSUMED STATES FOR PURPOSE OF PREPARING KEY.—21,2.<br />
REMARKS.—The compound falcigers <strong>of</strong> E. langi resemble<br />
those <strong>of</strong> E. tubifex, but compound spinigers are absent in E.<br />
langi. The aciculae and subacicular hooks are amber colored,<br />
ra<strong>the</strong>r than clear yellow; by definition this places <strong>the</strong> species in<br />
B-l as defined by Fauchald (1970). It is listed with similar<br />
species in Table 24 and 26.<br />
<strong>Eunice</strong> langi was considered synonymous with E. savignyi<br />
by Hartman (1956); it differs by having considerably darker<br />
subacicular hooks and acicula. In addition <strong>the</strong> latter have<br />
branchiae from setiger 4 through setiger 45 with a maximum <strong>of</strong><br />
12 filaments; E. langi has branchiae from setiger 3 through<br />
setiger 39 with a maximum <strong>of</strong> eight filaments. Due to <strong>the</strong><br />
indeterminate status <strong>of</strong> E. savignyi, o<strong>the</strong>r differences or<br />
similarities cannot be detailed at this time.<br />
None <strong>of</strong> <strong>the</strong> character states listed in Table 26 are unique to<br />
this species. Among species listed in Table 24, it is<br />
characterized first and foremost by <strong>the</strong> very poor development<br />
<strong>of</strong> <strong>the</strong> head <strong>of</strong> <strong>the</strong> compound falcigers and <strong>the</strong> sharply pointed,<br />
but not mucronate, hood <strong>of</strong> <strong>the</strong>se hooks.<br />
100. <strong>Eunice</strong> laticeps Ehlers, 1868<br />
FIGURE 62; TABLES 27, 30<br />
<strong>Eunice</strong> tentaculata Quatrefages, 1866:317-318.<br />
<strong>Eunice</strong> laliceps Ehlers, 1868:312.—Fauchald, 1986:251-252, figs. 46-50.<br />
MATERIAL EXAMINED.—Holotype <strong>of</strong> E. tentaculata Quatrefages,<br />
MNHN, Paris, A.l(R.)-1868-no. 52b. Port Western,<br />
Australia, coll. Quoy and Gaimard, 1839.<br />
COMMENTS ON MATERIAL EXAMINED.—The holotype <strong>of</strong> E.