A Review of the Genus Eunice - Smithsonian Institution Libraries

174 SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY
mm long; the current specimen, labeled as type, is larger, as
indicated below. It is unclear how this discrepancy has arisen:
it is here assumed that a simple lapsus has taken place and that
specimen here described in fact is the holotype.
DESCRIPTION.—Holotype incomplete, of unknown sex, with
138 setigers; length 30 mm; maximal width 2 mm at setiger 10;
length through setiger 10, 3.5 mm.
Prostomium (Figure 56j) distinctly shorter and narrower than
peristomium, as deep as x li of peristomium. Prostomial lobes
frontally rounded, somewhat flattened dorsally; median sulcus
very deep. Eyes behind bases of A-I, kidney-shaped, black.
Antennae in horseshoe, evenly spaced, similar in thickness.
Ceratophores ring-shaped in all antennae, without articulations.
Ceratostyles medially inflated with irregular notches, but
without articulations. A-I to middle of anterior peristomial ring;
A-II and A-III to posterior end of peristomium. Peristomium
longer than prostomium, cylindrical. Separation between rings
distinct on all sides; anterior ring ~ 2 h of total peristomial
length. Peristomial cirri barely to front edge of posterior
peristomial ring, digitiform, without articulations.
Maxillary formula 1+1,5+5, 7+0,6+9, and presumably 1+1;
Mx V apparently absent in preparation (see comment below).
Distal tooth of both Mx II with small subdistal notches,
indicating possible presence of reduced sixth tooth.
Branchiae absent.
Anterior neuropodial acicular lobes short, distally rounded,
becoming sessile in median and posterior setigers (Figure 56n);
aciculae emerging above midline. Pre- and postsetal lobes low,
transverse folds. Anterior ventral cirri digitiform, becoming
basally inflated by setiger 10. Ventral cirri retaining inflated
bases through rest of body, but inflated bases increasingly
retracted into body wall. Narrow tips short and button-shaped
in median setigers, becoming distinct and tapering in far
posterior setigers. Anterior notopodial cirri tapering and about
twice as long as ventral cirri, increasing in length through
median setigers. Notopodial cirri reduced in length from about
setiger 50, becoming short and tapering in far posterior setigers,
located well above neuropodia on body wall. Notopodial cirri
without articulations.
Pectinate setae (Figure 56m) tapering, flat. Both marginal
teeth longer than other teeth; -10 teeth present. Shafts of
compound falcigers (Figure 561) slender, distally strongly
inflated, marginally strongly serrated; distinct beak present.
Appendages very short, with nearly parallel sides; heads small,
bidentate. Both teeth similar in size. Proximal teeth slender,
tapering, directed laterally. Distal teeth tapering, directed
laterally. Guards symmetrically rounded, marginally serrated;
mucros absent. Pseudocompound falcigers and compound
spinigers absent. Aciculae (Figure 56n) single, with black core
and clear sheath, thick, distally tapering, straight; crosssections
round. Subacicular hooks (Figure 56k) with black core
and clear, nearly translucent heads, bidentate. Hooks first
present from setiger 50, present in all setigers thereafter, always
single (except for replacements). Hooks with distinctly bent
necks; heads distinct. Proximal teeth larger than distal teeth,
curved, directed laterally. Distal teeth triangular, tapering,
directed distally.
UNKNOWN MORPHOLOGICAL FEATURES.—Relationship between
Mx III and left Mx IV; pygidium and anal cirri.
EXPECTED STATES OF SELECTED UNKNOWN FEATURES.—
Mx III short and forming part of distal arc with left Mx IV.
CHARACTERS USED IN PREPARATION OF KEY NOT
SCORED.—Inappropriate Characters: 22, 31-42, 56, 60.
Unknown Characters: 1, 2.
ASSUMED STATES FOR PURPOSE OF PREPARING KEY.—
None.
REMARKS.—Eunice imogena is compared to similar species
in Tables 33 and 40. It is unique among the abranchiate species
in that subacicular hooks are first present from sctigcr 50; in the
other species, subacicular hooks arc present from sctigcrs
16-31.
The maxillary formula was given by Monro (1924:62) in the
following words: "formula as follows: 4-4:7+57-9." The
illustration given by Monro (1924, fig. 24) shows the formula
to be 1+1, 5+5, 7+0, 6+9. The jaws arc now missing; Monro's
illustration suggests a less aberrant formula than the text and is
here assumed to approximate the condition in the species.
89. Eunice impexa Grube, 1878
FIGURE 57i-m; TABLES 52,53
Eunice impexa Grube, 1878b: 159-160. pi. 9: Tig. 6.
MATERIAL EXAMINED.—Holotype, ZMB 4001, Cape of
Lapinig, Philippines, coll. Semper.
COMMENTS ON MATERIAL EXAMINED.—Four parapodia
removed from the holotype were also available; two of these
were previously examined by Grube, the other two by Augener.
DESCRIPTION.—Holotype incomplete, of unknown sex, with
101 setigers; length 63 mm; maximal width 5.5 mm at setiger
10; length through setiger 10, 11 mm.
Prostomium (Figure 57a) distinctly shorter than peristomium,
distinctly narrower than peristomium, less than x li as
deep as peristomium. Prostomial lobes frontally rounded,
dorsally flattened; median sulcus shallow. Eyes present
posterior to bases of A-I, faded purple. Antennae in horseshoe,
evenly spaced, similar in thickness. Ceratophores ring-shaped
in all antennae, without articulations. Ceratostyles tapering,
with 1 or 2 irregular, cylindrical articulations. A-I to anterior
peristomial ring; A-II to posterior peristomial ring; A-III to
setiger 1. Peristomium massive, expanded anteriorly due to
eversion of jaws; muscular lower lip present. Separation
between rings distinct ventrally and visible dorsally; anterior
ring 3 /4 of total peristomial length. Peristomial cirri to middle of
anterior peristomial ring, digitiform, without articulations.
Maxillary formula 1+1, 5+4, 5+0, 2+5, and 1+1. Mx III
short, forming a distal arc with very short left Mx IV and MX
V. Mx VI absent.